to accommodate the available evidence on site 2 substrates , it is concluded that the substrate hydrogens are first transferred to ubiquinone , 2 h+ per 2e then appear in the medium by protolytic dehydrogenation of a species of ubiquinol or ubiquinol-protein having the appropriate sidedness ( designated site 2a ) , and the other 2 h+ are translocated from the matrix to the medium on passage of 2e- through the cytochrome b x c1 complex ( designated site 2b ) .
C0020275
hydrogens
C0011744
2 h