experiments with analogues of the uncoupler pentachlorophenol , all of which are weaker acids than pentachlorophenol , indicated that the anionic form of the uncoupler is the potent form and we propose that it binds to a certain membrane protein to cause release into the cytoplasm of the substance ( ion , metabolite or protein ) that controls tumbling frequency .
C0003075
anionic
C0022023
ion