in doubtful cases their ne differentiation becomes established by a characteristic silver affinity , by the ultrastructurally observed presence of characteristic `` '' endocrine '' '' secretion granules , and by immunohistochemically detectable occurrence of `` '' pan-ne markers '' '' ( neuron-specific enolase , chromogranins , and synaptophysin ) , biogenic amines ( mainly serotonin ) , and neurohormonal peptides .
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