the extent of adp-ribosylation of histone h1 was also demonstrated to retard the protein 's mobility in acid-urea , acid-urea-triton , and lithium dodecyl sulfate polyacrylamide gels and to consist of at least 12 distinct adp-ribosylated species extractable in all nucleosome complexities studied .
C0019644
histone h1
C0028623
nucleosome