seed shriveling in the man-made intergeneric hybrid , triticale ( x triticosecale wittmack ) appears to be related to increased activity of endosperm acid phosphatases including para-nitrophenyl phosphatase , atpase , adpase , phosphatidic phosphatase , and glucose-1-phosphatase that occur specifically at later stages of seed development .
C0028189
para-nitrophenyl phosphatase
C0031678
phosphatase