the amino acids in subsites p3 ( pro , ala , leu , or asn ) , p2 ( gln , leu , hyp , arg , asp , or val ) , p1 ' ( ile or leu ) , and p4 ' ( gln , thr , his , ala , or pro ) all influence the hydrolysis rates .
C0002520
amino acids
C0017797
gln