the purified enzyme split amide and ester linkages in l-pyroglutamyl derivatives of l-alanine , beta-naphthylamine , alpha-naphthol , and 4-methylumbelliferone , but was completely inert towards various peptides and esters used as substrates for usual amino- and carboxy-peptidases , and for endopeptidases such as trypsin , subtilisin and alpha-chymotrypsin .
C0014442
enzyme
C0030946
endopeptidases