acisoma is a small genus of dragonflies in the family libellulidae .

it contains six species : acisoma ascalaphoides rambur , 1842 – littoral pintail acisoma attenboroughi mens , schütte , stokvis & dijkstra , 2016 – attenborough 's pintail acisoma inflatum selys , 1882 – stout pintail acisoma panorpoides rambur , 1842 – asian pintail , grizzled pintail acisoma trifidum kirby , 1889 – pied pintail , ivory pintail acisoma variegatum kirby , 1898 – slender pintail

acisoma

acisoma <EOT> dragonflies & damselflies of thailand : 104 . acisoma panorpoides panorpoides ( rambur , 1842 )

kento furui added an unknown common name in an unknown language to " acisoma " .

the type locality for acisoma panorpoides ascalaphoides is madagascar . it is not currently certain that the populations of madagascar and mainland africa are similar ( k . schütte in dijkstra and boudot 2010 ) , so it is likely that the present subspecific name will have to change in the future for africa . this species previously appeared on the iucn red list as acisoma panorpoides ssp . ascalaphoides but it has now been promoted to acisoma ascalaphoides .

acisoma panorpoides perches inconspicuously . it tends to be more common on short ranked grasses near water bodies . it is mainly associated with eutrophic still waters ( d . h . murphy pers . comm . ) .

justification : acisoma trifidum is listed as least concern because of its wide distribution , as there are no major threats and as it is unlikely to be declining fast enough to qualify for listing in a threatened category .

justification : acisoma variegatum occurs from southern africa up to ethiopia . as this species is common in the southern part of its range ( south africa , mozambique , zimbabwe ) , it is assessed as least concern .

acisoma variegatum is common in southern africa and the distribution extends up to ethiopia . it was formerly confused with a . inflatum and both were once listed under a . panorpoides . due to this confusion , the extent of occurrence ( eoo ) of a . variegatum and a . inflatum requires further study .

formerly acisoma panorpoides was thought to occur in africa , madagascar and asia . recent taxonomic and genetic studies showed , that in asia the true a . panorpoides occurs , while two different species each occur in madagascar and continental africa . the latter are separated in a . variegatum and in a . inflatum , both of them being described long ago and neglected since ( dijkstra and clausnitzer 2014 ) .

the dragonfly genus acisoma is revised based on adult male morphology and coi sequence data . six species are recognised , including the new species a . attenboroughi sp . nov . diagnoses and a key to males of all species and illustrations of all relevant characters are provided . a . inflatum , a . variegatum and a . trifidum are confined to continental africa , while a . panorpoides is restricted to asia . a . ascalaphoides is known only from threatened littoral forest fragments on the east coast of madagascar , while a . attenboroughi is widespread across the island . the new species honours sir david attenborough on his 90 th birthday .

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thank you for taking the time to provide feedback on the iucn red list of threatened species website , we are grateful for your input .

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schorr , m . and paulson , d . 2016 . world odonata list . revision 21 june 2016 . tacoma , washington , usa available at : urltoken .

endangered b2ab ( ii , iii , iv , v ) ver 3 . 1

justification : this species is endemic to madagascar where it known from three littoral forest sites : two in the south east and one from the northern east coast . it has an extent of occurrence ( eoo ) of 5 , 884 km² and an area of occupancy ( aoo ) of 16 km² . the species experiences continuing decline in its habitat quality and extent due to ongoing deforestation in the north east and mining activities in the south east parts of its range . based on these threats 2 - 3 locations have been identified . therefore , the species is assessed as endangered .

the species is endemic to madagascar where it is known from one locality in the north east coast in voloina and two localities in mandena and sainte luce in the south east part of the island .

known only from littoral forest fragments at the southern ( tolagnaro , formerly fort dauphin ) and northern ( voloina ) ends of madagascar’s east coast . only about 10 % of the original cover of these forests remains , all in small patches of which only 13 % are protected ( consiglio et al . 2006 ) . the species may occur elsewhere along the coast , but as it seems restricted to this habitat , could well be under threat ( schütte &

razafindraibe 2007 ) . the larvae are possibly adapted to more acidic water than a . attenboroughi sp . nov .

the species is threatened by deforestation in the north and mining activities in the south within the mining offset / conservation sites .

no conservation actions are known . the species doesn ' t occur within protected areas in the north . in the south the species is found within the mining offset conservation sites .

dijkstra , k . d . and clausnitzer , v . 2014 . the dragonflies and damselflies of eastern africa : handbook for all odonata from sudan to zimbabwe . rmca .

no information on the population size is available , but this species is common in southern africa ( south africa , mozambique , zimbabwe ) .

this species occurs in swampy and well vegetated habitats along lakes , pools and slow parts of rivers and streams in savannah and woodland .

drainage , destruction and pollution of swampy habitats for construction of human settlements are the main threats to this species . swamps close to settlements suffer particularly from pollution due to sewage and to illegal rubbish damping .

the species is widespread and therefore , no conservation actions are necessary . however , it is necessary to study the extent of occurrence ( eoo ) of this species due to changes in taxonomy .

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this dragonfly is easily recognised by its uniquely shaped abdomen , which tapers markedly from segments five to seven to form a narrow tip . the male has light blue eyes and body , with dark markings on the thorax and abdomen . the female is similarly shaped and patterned , but with greenish yellow body . in males , the hindwing is 20 to 22 mm in length and the total body leangth ranges from 27 to 29 mm .

commonly seen in disturbed habitats and grassy swamps throughout open country in singapore , including open areas in central catchment and bukit timah nature reserve and freshwater ponds in sungei buloh wetland reserve .

when a female oviposits , the male guards her closely , seldom straying more than 40 cm from her .

tang , h . b . , l . k . wang & m . hämäläinen , 2010 . a photographic guide to the dragonflies of singapore . raffles museum of biodiversity research , national university of singapore , singapore . 222 pp .

raffles museum of biodiversity research rmbr has its origins in the raffles museum which was founded in 1849 . established on…

developed by moving mouse © 2018 lee kong chian natural history museum . all rights reserved . terms of use .

joshi , s . , p . koparde , p . dawn , p . roy , and k . kunte ( eds . ) .

copyright © 2018 , all rights reserved . national centre for biological sciences ( ncbs ) holds copyright for all the original material and compilations on this website , although contributing writers and photographers may hold copyright for their material as cited . material from this website can be used freely for educational , basic research and conservation purposes , provided that this website is acknowledged and properly cited as the source . contact us to obtain prior permission for any other use , including for large data downloads and collaborative research .

lotte p . mens naturalis biodiversity center , p . o . box 9517 , 2300 ra leiden , the netherlands .

kai schütte universität hamburg , zoological institute , animal ecology and conservation and center of natural history , zoological museum , martin - luther - king - platz 3 , 20146 hamburg , germany .

frank r . stokvis naturalis biodiversity center , p . o . box 9517 , 2300 ra leiden , the netherlands .

klaas - douwe b . dijkstra naturalis biodiversity center , p . o . box 9517 , 2300 ra leiden , the netherlands . department of conservation ecology and entomology , university of stellenbosch , south africa

lotte p . mens , kai schütte , frank r . stokvis , klaas - douwe b . dijkstra

from a distance . it ' s only when you get close ( which is actually quite difficult ) that you notice its big bulbous abdomen .

the male is quite stunning , with its bright blue eyes , and blue and black markings to the thorax and abdomen . it almost has a marble effect . here you can see just how bulbous the abdomen really is . s9 - 10 are black and the caudal appendages are white . the young male is yellow , as the female .

the female is very similar to the male , except it is yellow in colour instead of blue and the caudal appendages are wider apart ( usually the only way i can identify the female and young males ) .

here is a young female , i managed to capture at nam nao recently .

you can see this species throughout the country and it seems most common in the cool season ( dec - mar ) . it is very difficult to get close to and flies away at the slightest movement . the females are slightly more forgiving than the males , but it ' s worth the effort , because it really is a good - looking dragonfly .

i am too lazy to write about my past , but i now love photographing dragonflies , manchester city football club , fishing and , of course , my girlfriend .

98 . ischnura sp . ( rufostigma selys , 1876 - group ) . . .

number : 186 family : libellulidae genus : nannophya species : nannophya pygmaea common name ( s ) : the scarlet dwarf . . .

number : 182 family : coenagrionidae genus : ceriagrion species : ceriagrion malaisei common name ( s ) : n / a synonyms : . . .

location : phu kao - phu phan kham national park , khon kaen date : saturday 28th may , 2016 habitat : lowland , shallow lake on the edg . . .

number : 176 family : lestidae genus : platylestes species : platylestes platystylus common name ( s ) : n / a synonyms : n / a . . .

location : phu khieo wildlife sanctuary , chaiyaphum date : saturday , 12th november , 2016 habitat : mid - to upland forested ponds . . .

number : 175 family : libellulidae genus : lyriothemis species : lyriothemis sp . common name ( s ) : n / a synonyms : n / a ha . . .

location 1 : tat fa and pha ing waterfalls , tat ton national park , chaiyaphum date : saturday 26th march , 2016 habitat : lowlands ( a . s . l . . . .

number : 189 family : libellulidae genus : amphithemis species : amphithemis curvistyla common name ( s ) : n / a synonyms : . . .

number : 185 family : coenagrionidae genus : ceriagrion species : ceriagrion pallidum common name ( s ) : n / a syn . . .

number : 57 family : libellulidae genus : trithemis species : trithemis aurora common name ( s ) : crimson marsh glider , crimson dropwing , . . .

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is widespread in africa ( except dense rain forest ) , southern europe , middle east , southern asia , and indian ocean islands .

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peripsocus milleri is a species of psocoptera from peripsocidae family that can be found in great britain and ireland .

they can also be found on azores and canary islands , belgium , croatia , france , italy , and spain .

the species are brown coloured .

peripsocus milleri

peripsocus milleri <EOT> lienhard & courtenay smithers . 2002 . psocoptera ( insecta ) world catalogue and bibliography > > note : catalog > > peripsocus milleri

thornton & wong . 1968 . pacific insects monographs 19 : 129 > > note : hawaii , new zealand > > peripsocus nitens urn : lsid : psocodea . speciesfile . org : taxonname : 5577

new , t . r . 1973 , " new species and records of peripsocus hagen ( psocoptera , peripsocidae ) from southeast australia " , journal of the australian entomological society , vol . 12 , pp . 340 - 346 16 figs

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confused by a class within a class or an order within an order ? please see our brief essay .

to cite this page : myers , p . , r . espinosa , c . s . parr , t . jones , g . s . hammond , and t . a . dewey . 2018 . the animal diversity web ( online ) . accessed at https : / / animaldiversity . org .

disclaimer : the animal diversity web is an educational resource written largely by and for college students . adw doesn ' t cover all species in the world , nor does it include all the latest scientific information about organisms we describe . though we edit our accounts for accuracy , we cannot guarantee all information in those accounts . while adw staff and contributors provide references to books and websites that we believe are reputable , we cannot necessarily endorse the contents of references beyond our control .

this material is based upon work supported by the national science foundation grants drl 0089283 , drl 0628151 , due 0633095 , drl 0918590 , and due 1122742 . additional support has come from the marisla foundation , um college of literature , science , and the arts , museum of zoology , and information and technology services .

if you have images for this taxon that you would like to share with atlas of living australia , please upload using the upload tools .

edwards , b . a . b . 1950 , " a study of the tasmanian psocoptera with descriptions of new species " , papers and proceedings of the royal society of tasmania , vol . 1949 , pp . 93 - 134 117 figs

tillyard , r . j . 1923 , " a monograph of the psocoptera , or copeognatha , of new zealand " , transactions of the new zealand institute , vol . 54 , pp . 170 - 196 20 figs pl . 18

urn : lsid : biodiversity . org . au : afd . taxon : a28da8d9 - 9346 - 4a05 - b504 - 526cf30901a4

urn : lsid : biodiversity . org . au : afd . taxon : ecbc232f - be9a - 4fac - a417 - ced19e9d7b7b

urn : lsid : biodiversity . org . au : afd . taxon : 43fbc356 - 79a9 - 4533 - 9c8a - 1cddf50b0d72

urn : lsid : biodiversity . org . au : afd . name : 349041

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paulo a . v . borges , 1 clara gaspar , 1 luís carlos fonseca crespo , 1 , 2 françois rigal , 3 , 1 pedro cardoso , 4 , 1 fernando pereira , 1 carla rego , 1 isabel r . amorim , 1 catarina melo , 1 carlos aguiar , 5 genage andré , 5 enésima p . mendonça , 1 sérvio ribeiro , 6 , 1 joaquín hortal , 7 , 1 ana m . c . santos , 7 , 1 luís barcelos , 1 henrik enghoff , 8 volker mahnert , 9 margarida t . pita , 10 jordi ribes , 11 arturo baz , 12 antónio b . sousa , 13 virgílio vieira , 1 , 14 jörg wunderlich , 15 , 15 aristeidis parmakelis , 16 , 1 robert j . whittaker , 17 josé alberto quartau , 5 artur r . m . serrano , 5 and kostas a . triantis 16 , 1

4 finnish museum of natural history , university of helsinki , pohjoinen rautatiekatu 13 , p . o . box 17 , 00014 , helsinki , finland

12 dep . de ciencias de la vida . universidad de alcalá , 28871 alcalá de henares , madrid , spain

14 departamento de biologia , universidade dos açores , apartado 1422 , 9501 - 301 , ponta delgada , s . miguel , azores , portugal

this is an open access article distributed under the terms of the creative commons attribution license 4 . 0 ( cc - by ) , which permits unrestricted use , distribution , and reproduction in any medium , provided the original author and source are credited .

in this contribution we present detailed distribution and abundance data for arthropod species identified during the bala – b iodiversity of a rthropods from the l aurisilva of the a zores ( 1999 - 2004 ) and bala2 projects ( 2010 - 2011 ) from 18 native forest fragments in seven of the nine azorean islands ( all excluding graciosa and corvo islands , which have no native forest left ) .

azores ; terrestrial arthropods ; bala project ; laurissilva forest ; linnean , wallacean and prestonian shortfalls .

in 1999 a group of researchers from the university of the azores and the university of lisbon started a long - term ( 1999 - 2004 ) standardized sampling program to inventory the arthropod biodiversity in native forest remnants of the azores - the bala i project – b iodiversity of a rthropods from the l aurisilva of the a zores ( borges et al . 2000 , borges et al . 2005a , borges et al . 2011 , ribeiro et al . 2005 , gaspar et al . 2008 ) . more recently , this project was extended by researchers from the universities of the azores , athens and oxford , by surveying part of the same native forest plots almost 10 years later - bala ii project ( 2010 - 2011 ) .

during this period , two complete checklists of azorean arthropod fauna were produced ( borges et al . 2005b , borges et al . 2010 ) , which included the distribution of each species per island . in this paper we compile and synthesize the faunistic results of both bala projects , highlighting novel distribution records and presenting not only detailed distribution but also abundance data for each species , adding taxonomical and biogeographical information whenever possible . finally , we provide a general and updated overview on the diversity of the azorean arthropods .

the remote azores archipelago extends for 615 km in the north atlantic ocean ( 37 - 40 °n , 25 - 31 °w ) , 1584 km to the east ( southern europe ) and 2150 km to the west ( northern america ) of the nearest mainland . it comprises nine main islands and some small islets , all of volcanic origin , and is located at the triple junction of the eurasian , african and american tectonic plates . the nine islands are divided into three groups : the western group ( corvo and flores isls . ) , the central group ( faial , pico , graciosa , são jorge and terceira isls . ) , and the eastern group ( são miguel and santa maria isls ) ( fig .

) . the climate is temperate and oceanic , strongly influenced by the ocean and island topography , which together produce high relative atmospheric humidity , above 95 % on average on native forests .

location of the azores and of native forest fragments in the archipelago . codes for forest fragments as in table

) . graciosa and corvo islands were excluded as they no longer present native forest . human settlement in the azores lead to considerable native forest destruction which has left the entire archipelago with little over 2 % of the original forest cover . during the summer ( june to september ) 150 m long and 5 m wide transects were set up in 100 sites from 1999 to 2004 (

project ; 15 native forest fragments ) . along each transect , arthropods from the soil ( mainly epigean ) and herbaceous vegetation were surveyed with pitfall traps , while arthropods from woody plants were sampled using a beating tray . pitfall traps consisted of plastic cups with 4 . 2 cm diameter and 7 . 8 cm height . thirty pitfall traps were set up per transect . half of the traps were filled with a non - attractive ethylene glycol preservative solution ( antifreeze solution ) , and the remaining with a general attractive solution , a modified version of turquin (

) prepared mainly with dark beer and preservative agents . a few drops of dishwashing liquid were added to both solutions to reduce surface tension . traps were sunk in the soil ( cup rim at surface level ) every 5 m along the transects , those filled with turquin alternating with traps containing antifreeze solution . traps were protected from rain using a plastic plate , placed about 5 cm above surface level and fixed to the ground by two pieces of wire . accidental collection of small vertebrates and damage by rodents was prevented using a piece of plastic mesh placed on top of the trap and fixed to the ground by pieces of wire . the traps remained active in the field for two weeks .

main characteristics of the azorean islands ( bold ) and native forest fragments sampled from 1999 to 2011 , including area ( hectares ) , highest point ( altitude in metres ) , distance to the nearest island / fragment ( isolation in kilometres ) and the oldest geological age of emerged substrate ( million years bp ) ( adapted from gaspar et al . 2008 ) .

canopy sampling was conducted during the trapping period , when the vegetation was dry . a 5 m wide square was established every 15 m ( total of 10 squares per transect ) . two woody plant specimens of the most abundant species ( up to three species when available ) were sampled in each square . for each selected plant , a branch was chosen at random and a beating tray placed beneath . the tray consisted of a 1 m wide and 60 cm deep cloth inverted pyramid , with a plastic bag at the vertex . five beatings were made using a stick for each plant individual sampled .

all specimens are deposited in the entomological collection dalberto teixeira pombo at the university of the azores ( portugal ) , under the curation of paulo a . v . borges ( pborges @ urltoken ) .

in this contribution we list the 286 species for which we obtained an identification . the new records for each island are marked with * . for this list two families of coleoptera were not considered since they will be presented elsewhere , staphylinidae ( borges et al . in prep . ) and zopheridae ( borges et al . 2016 ) . for detailed maps on the distribution of these species in azores consult the azores bioportal .

all specimens were assigned a site code composed of several letters and numbers that read as follows ( see suppl . material 1 for complete data ) . detailed metadata is given in suppl . material 2 ) :

iv ) the next letter refers to the sampling technique : p - pitfall , b - canopy beating ; for pitfall samples ( p ) tu – turquin and et – ethylene glycol ; for canopy samples ( b ) the next two letters refer to the plant sampled : ca = calluna vulgaris , cl = clethra arborea , er = erica azorica , fr = frangula azorica , il = ilex perado azorica , ju = juniperus brevifolia , la = laurus azorica , mc = morella faya , ms = myrsine africana , pi = picconia azorica , pt = pittosporum undulatum , va = vaccinium cylindraceum .

for the geographical location of transects within reserves ( utm coordinates ) see suppl . material 3 .

accumulation curves were obtained using the software “ species diversity and richness ” v . 4 .

the ultimate goal of biodiversity assessments is documenting all species inhabiting a region . however , this has often proven impossible to achieve given the unfeasibility of collecting every single species that exists in a study area . this study focuses on the terrestrial arthropod diversity of the azores and encompasses most orders of the phylum

, we excluded 47 % of the archipelago ' s species pool . yet , this study added 10 endemic and at least 16 other species , mostly exotics , to the known azorean arthropod fauna . more will be added soon after the on - going revision of

, in press ) . overall , at least 26 species that occur in native forests were added to the azorean arthropod fauna list . the new 346 taxonomic records provided by this study ( see suppl . material

) . 164 species were found in new islands , with an average of two islands per species . for 82 of those species only one new island was added to their known distribution contrasting with 27 species for which four or more islands were added ( fig .

species richness for the azores archipelago and each island . total currently known species , the number of species surveyed during this study and those that represent new records are presented .

total species and subspecies records for the azores , new species and subspecies records during this study and increment for the most speciose classes and orders . values for all islands are added , so richness may be up to 7 times higher than the archipelago ' s richness ( as 7 islands were surveyed ) . ( * ) the coleoptera families staphylinidae and zopheridae were not considered ( see text ) .

the number of species identified for each of the 18 native forest fragments surveyed is shown in fig .

. the fragment with the highest species diversity is serra de santa bárbara in terceira island ( s = 124 ) , which is also the larger native forest area in the azores . remarkably , one of the smallest fragments , pico alto in santa maria island , is the second most diverse ( s = 121 ) .

bala2 samples only added 4 % of species to the previous bala survey ( fig .

) . interestingly , 59 samples collected in the first two years of survey ( 1999 and 2000 ) provided about 81 % of the total species recorded in this study .

species accumulation curve for the 286 species of arthropods collected in 152 pitfall and beating samples between 1999 and 2011 . the solid line corresponds to the chronological sample sequence and the dotted line is a randomized curve ( 1000 runs ) . samples to the left of the vertical line were collected in bala1 and to the right in bala2 .

a total of 163744 individuals were identified as belonging to the 286 species ( see suppl . material

for the complete list of abundance per species ) . the ten most abundant species ( fig .

) accommodate 56 % of the total number of individuals and include mostly indigenous species ( endemic or native non - endemic ) . the single introduced species is the millipede

) that are mostly soil epigean species , the other seven species live preferentially in the canopies of azorean endemic trees . the moth

the ten most abundant species in the database . end - endemic from azores ; nat - native non - endemic species ; intr - species introduced in the archipelago .

the opilion leiobunum blackwalli ( credit : paulo a . v . borges ) .

the increase in the number of islands from where each species is known and the distribution increase for many species within each island shows the importance of regional standardized surveys , which provided a major improvement in the knowledge of the distribution of arthropod species in the native forests of the azores .

the fact that most diversity was captured during the first two years of the project reflects the importance of sampling a wide geographic range covering all the islands and the maximum number of sites . increasing the number of samples per fragment ( sampling performed in 2004 ) or replicating the sampling at a different time ( 29 sites in 2010 to 2011 ;

project ) had a lesser impact in increasing our knowledge about biodiversity ( fig .

a ) expanding the standardized survey of azorean arthropods to other habitat types , mostly man - modified , an already on - going task for some of the islands ( see e . g . cardoso et al . 2009 , meijer et al . 2011 , cardoso et al . 2013 , florencio et al . 2013 , santos et al . 2010 ) ;

b ) selecting study areas along a comprehensive environmental gradient where an optimal sampling strategy will be applied in order to sample the entire arthropod communities ( all taxa biodiversity inventory - atbi ) . atbis are intensive sampling efforts to identify and record all living species that exist within a given area and simultaneously create a common and standardized biodiversity database ( lawton and gaston 2001 ) ;

c ) finishing the identification of many morphospecies . good progress has been made with staphylinidade ( borges et al . in prep . ) , but other taxa need further effort to reach proper identification ;

d ) increase sampling and update the current list of azorean hymenoptera and diptera , which is clearly incomplete ( borges et al . 2010 ) . the shortage of taxonomists who can adequately identify species ( i . e . the so - called taxonomic impediment ) has prevented advances in the knowledge for many diverse groups in the azores , including these two .

e ) contributing to the validation and updating of the pan - european checklists programs , including fauna europaea ( jong et al . 2014 ) and pesi ( jong et al . 2015 ) allowing a more general evaluation and comparison of species distributions and statuses .

this study advances the knowledge on the unique arthropod biodiversity of the azores , but at the same time highlights the need for further surveys . we strongly believe that the bala project will stimulate further research and conservation actions towards the preservation of azorean biodiversity . furthermore , we hope that all the taxa yet to be identified will entice taxonomist to join us in the endeavour of cataloguing all terrestrial arthropods of the most remote of the macaronesian archipelagos , the azores . the ongoing longterm research projects in azores and the recent creation of the e - repository islandlab will create new opportunities for biodiversity studies in azores .

brief description : detailed data on the occurrences and abundances of the studied species . data on species abundance in each individual sample ( pitfall trap or canopy beating ) for the 152 transects in eighteen protected areas and seven azorean islands .

brief description : utm coordinates ( regions 25s for flores and 26s for all other islands ) , altitude ( meters ) and supporting project of the studied transects in the azores . transect code according to island , reserve and transect number ( see text )

pavb , arms , jq and kat conceived the ideas ; pavb , cg , lc , fr , pc , fp , cr , ira , cm , ca , ga , spr , jh , amcs , abs , jw , jaq , arms and kat obtained samples ; pavb , cg , em and lb prepared the databases ; pavb , lc , pc , he , fi , vm , mtp , jr , ab , abs , rzs , vv , jw , jaq and arms performed taxonomic work and identified species ; pavb led the writing with substantial input from the other authors .

blas m , borges pav . a new species of catops ( coleoptera : leiodidae , cholevinae ) from the azores with remarks on the macaronesian fauna .

borges p . a . v . , serrano a . r . , quartau j . a . ranking the azorean natural forest reserves for conservation using their endemic arthropods .

erichson , 1845 ( coleoptera : zopheridae ) : an integrative taxonomic approach with description of four new species .

borges p . a . v . , gaspar c . s . , santos a . m . c , ribeiro s . p . , cardoso p . , triantis k . , amorim i . r . patterns of colonization and species distribution for azorean arthropods : evolution , diversity , rarity and extinction ; celebrating darwin : proceedings of the symposium " darwin ' s mistake and what we are doing to correct it ; ponta delgada . açoreana ; 2011 . 30 .

borges p . a . v . , vieira v . , amorim i . r . , bicudo n . , fritzén n . , gaspar c . , heleno r . , hortal j . , lissner j . , logunov d . , machado a . , marcelino j . , meijer s . s . , melo c . , mendonça e . p . , moniz j . , pereira f . , santos a . s . , simões a . m . , torrão e . list of arthropods ( arthropoda ) in : borges p . a . v , costa a . , cunha r . , gabriel r . , gonçalves v . , martins a . f . , melo i , parente m . , raposeiro p . , rodrigues p . , santos r . s . , silva l . , vieira p . , vieira v . , editors .

borges p . a . v . , aguiar c . , amaral j . , amorim i . r . , andré g . , arraiol a . , baz a . , dinis f . , enghoff h . , gaspar c . , ilharco f . , mahnert v . , melo c . , pereira f . , quartau j . a . , ribeiro s . p . , ribes j . , serrano a . r . m . , sousa a . b . , strassen r . z . , vieira l . , vieira v . , vitorino a . , wunderlich j . ranking protected areas in the azores using standardised sampling of soil epigean arthropods .

borges p . a . v . , vieira v . , dinis f . , jarroca s . , aguiar c . , amaral j . , aarvik l . , ashmole p . , ashmole m . , amorim i . r . , andré g . , argente m . c . , arraiol a . , cabrera a . , diaz s . , enghoff h . , gaspar c . , mendonça e . p . , gisbert h . m . , gonçalves p . , lopes d . h . , melo c . , mota j . a . , oliveira o . , oromí p . , pereira f . , pombo d . t . , quartau j . a . , ribeiro s . p . , rodrigues a . c . , santos a . m . c . , serrano a . r . m . , simões . a . m . , , soares a . o . , sousa a . b . , vieira l . , vitorino a , wunderlich j . list of arthropods ( arthropoda ) in : borges p . a . v . , cunha r . , gabriel r . , martins a . m . f . , silva l . , vieira v . , editors .

a list of the terrestrial fauna ( mollusca and arthropoda ) and flora ( bryophyta , pteridophyta and spermatophyta ) from the azores .

direcção regional de ambiente & universidade dos açores ; horta , angra do heroísmo & ponta delgada : 2005 . 58 .

borges p . a . v . , serrano a . r . m . , amorim i . r . new species of cave - dwelling beetles ( coleoptera : carabidae : trechinae ) from the azores .

borges paulo a . v . , wunderlich joerg . spider biodiversity patterns and their conservation in the azorean archipelago , with descriptions of new species .

borges paulo a . v . , lobo jorge m . , azevedo eduardo b . , gaspar clara s . , melo catarina , nunes luis v . invasibility and species richness of island endemic arthropods : a general model of endemic vs . exotic species .

cardoso pedro , borges paulo a . v . , gaspar clara . biotic integrity of the arthropod communities in the natural forests of azores .

cardoso pedro , erwin terry l . , borges paulo a . v . , new tim r . the seven impediments in invertebrate conservation and how to overcome them .

cardoso p . , rigal f . , fattorini s . , terzopoulou s . , borges p . a . v . integrating landscape disturbance and indicator species in conservation studies .

cardoso pedro , aranda silvia c . , lobo jorge m . , dinis francisco , gaspar clara , borges paulo a . v . a spatial scale assessment of habitat effects on arthropod communities of an oceanic island .

crespo lc , bosmans r , cardoso p , borges pav . on the endemic spider species of the genus savigniorrhipis wunderlich , 1992 ( araneae : linyphiidae ) in the azores ( portugal ) , with description of a new species .

crespo lc , bosmans r , cardoso p , borges pav . on three endemic species of the linyphiid spider genus canariphantes wunderlich , 1992 ( araneae , linyphiidae ) from the azores archipelago .

florencio m . , cardoso p . , lobo j . m . , azevedo e . b . , borges p . a . v . arthropod assemblage homogenization in oceanic islands : the role of exotic and indigenous species under landscape disturbance .

florencio m , lobo jm , cardoso p , almeida - neto m , borges pav . the colonisation of exotic species does not have to trigger faunal homogenisation : lessons from the assembly patterns of arthropods on oceanic islands .

gaspar c . , borges p . a . v . , gaston k . j . diversity and distribution of arthropods in native forests of the azores archipelago .

gaspar clara , gaston kevin j . , borges paulo a . v . , cardoso pedro . selection of priority areas for arthropod conservation in the azores archipelago .

gaston kevin j , borges paulo a v , he fangliang , gaspar clara . abundance , spatial variance and occupancy : arthropod species distribution in the azores .

hortal j , borges pav , gaspar c . evaluating the performance of species richness estimators : sensitivity to sample grain size .

jong yde de , verbeek melina , michelsen verner , place bjørn per de , los wouter , steeman fedor , bailly nicolas , basire claire , chylarecki przemek , stloukal eduard , hagedorn gregor , wetzel florian , glöckler falko , kroupa alexander , korb günther , hoffmann anke , häuser christoph , kohlbecker andreas , müller andreas , güntsch anton , stoev pavel , penev lyubomir . fauna europaea – all european animal species on the web .

lawton j . h . , gaston k . j . indicator species . in : levin s . a . , editor .

lobo j . , borges p . a . v . the provisional status of arthropod inventories in the macaronesian islands . in : serrano a . r . m . , borges p . a . v . , boieiro m . , oromí p . , editors .

martín josé l . , cardoso pedro , arechavaleta manuel , borges paulo a . v . , faria bernardo f . , abreu cristina , aguiar antónio f . , carvalho josé a . , costa ana c . , cunha regina t . , fernandes francisco m . , gabriel rosalina , jardim roberto , lobo carlos , martins antónio m . f . , oliveira paulo , rodrigues pedro , silva luís , teixeira dinarte , amorim isabel r . , homem nídia , martins berta , martins mónica , mendonça enésima . using taxonomically unbiased criteria to prioritize resource allocation for oceanic island species conservation .

meijer seline s . , whittaker robert j . , borges paulo a . v . the effects of land - use change on arthropod richness and abundance on santa maria island ( azores ) : unmanaged plantations favour endemic beetles .

platia g . , borges p . a . v . description of a new species of athous and record of the female of a . azoricus platia & gudenzi from the azores ( coleoptera , elateridae )

ribeiro sérvio p . , borges paulo a . v . , gaspar clara , melo catarina , serrano artur r . m . , amaral joão , aguiar carlos , andré genage , quartau josé a . canopy insect herbivores in the azorean laurisilva forests : key host plant species in a highly generalist insect community .

ribes j . , borges p . a . v . a new subspecies of orthotylus junipericola linnavuori , 1965 ( heteroptera ; miridae ) from the azores .

rigal françois , whittaker robert j . , triantis kostas a . , borges paulo a . v . integration of non - indigenous species within the interspecific abundance–occupancy relationship .

santos a . m . c . , borges p . a . v , rodrigues a . c . , lopes d . j . h . lista de espécies de artrópodes associados a diferentes culturas frutícolas da ilha terceira ( açores )

terzopoulou sofia , rigal françois , whittaker robert j . , borges paulo a . v . , triantis kostas a . drivers of extinction : the case of azorean beetles .

triantis kostas a . , borges paulo a . v . , ladle richard j . , hortal joaquín , cardoso pedro , gaspar clara , dinis francisco , mendonça enésima , silveira lúcia m . a . , gabriel rosalina , melo catarina , santos ana m . c . , amorim isabel r . , ribeiro sérvio p . , serrano artur r . m . , quartau josé a . , whittaker robert j . extinction debt on oceanic islands .

turquin m . une biocenose cavernicole originale pour le bugey : le puits de rappe . comptes rendus 96e congresse naturel societes savantes , toulouse 1071 .


lachnocnema pseudobibulus is a butterfly in the lycaenidae family .

it is found in uganda , kenya , tanzania , the democratic republic of the congo , malawi and zimbabwe .

lachnocnema pseudobibulus

lachnocnema pseudobibulus <EOT> lachnocnema , commonly called woolly legs , is a genus of butterfly in the family lycaenidae found mainly in subsaharan africa .

libert , m . ( 1996a ) contribution a l ' etude des lycaenidae africains . revision du genre lachnocnema trimen , ( lepidoptera lycaenidae ) . lambillionea 96 ( 1 ) : 185 - 202 .

libert , m . 1996 contribution a l ' etude des lycaenidae africains - revision du genre lachnocnema trimen ( lepidopter lycaenidae ) . lambillionea 96 , 185 - 202 , 367 - 386 , 479 - 500 .

libert , m . ( 1996c ) contribution a l ' etude des lycaenidae africains : revision du genre lachnocnema trimen , troisieme partie . ( lepidoptera lycaenidae ) . lambillionea 96 ( 3 ) : 479 - 500 .

libert , m . ( 1996b ) contribution a l ' etude des lycaenidae africains : revision du genre lachnocnema trimen , deuxieme partie ( 1 ) . ( lepidoptera lycaenidae ) . lambillionea 96 ( 2 ) : 367 - 386 .

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an african genus to which many species have been added recently due to the revision of libert ( 1996 ) .

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vári , l . , kroon , d . m . , & krüger , m . 2002 . classification and checklist of the species of lepidoptera recorded in southern africa . simple solutions , chatswood australia .

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woodhall , steve . field guide to butterflies of south africa . cape town : struik publisher , 2005 .

seitz , a . die gross - schmetterlinge der erde 13 : die afrikanischen tagfalter . plate xiii 65 template : miletinae - stub


venericardia is a widely distributed genus of marine bivalve molluscs , in the family carditidae .

it is the type genus of subfamily venericardiinae .

the closely related purpurocardia was for long included here as a subgenus , but is increasingly considered distinct .

species of venericardia include : venericardia amabilis venericardia bimaculata venericardia ferruginea venericardia granulata † venericardia imbricata ( gmelin , 1791 ) † venericardia iheringi ( boehm , 1903 ) venericardia planicosta

venericardia

venericardia <EOT> species venericardia dilecta accepted as vimentum dilectum ( e . a . smith , 1885 )

species venericardia excelsior accepted as vimentum dilectum ( e . a . smith , 1885 )

species venericardia alaskana dall , 1903 accepted as cyclocardia crebricostata ( a . krause , 1885 )

variety venericardia borealis var . ovata riabinina , 1952 accepted as cyclocardia ovata ( riabinina , 1952 )

species venericardia flammea michelin , 1831 accepted as strophocardia megastropha ( j . e . gray , 1825 )

species venericardia nodulosa dall , 1919 accepted as cyclocardia bailyi ( j . q . burch , 1944 )

species venericardia tankervillii w . wood , 1828 accepted as cardiocardita tankervillii ( w . wood , 1828 )

species venericardia zelandica potiez & michaud , 1844 accepted as austrovenus stutchburyi ( w . wood , 1828 )

species venericardia africana bartsch , 1915 accepted as coripia elata ( g . b . sowerby iii , 1892 )

species venericardia reinga a . w . b . powell , 1933 accepted as purpurocardia reinga ( powell , 1933 )

species venericardia minima ( e . a . smith , 1904 ) accepted as coripia agulhasensis ( jaeckel & thiele , 1931 )

species venericardia kiiensis ( g . b . sowerby iii , 1913 ) accepted as megacardita ferruginosa ( adams & reeve , 1850 )

species venericardia elata ( g . b . sowerby iii , 1892 ) accepted as coripia elata ( g . b . sowerby iii , 1892 )

species venericardia megastropha j . e . gray , 1825 accepted as strophocardia megastropha ( j . e . gray , 1825 ) ( original combination )

species venericardia crassicostata g . b . sowerby i , 1825 accepted as cardites crassicostatus ( g . b . sowerby i , 1825 ) ( original combination )

species venericardia lilliei c . a . fleming , 1943 † accepted as purpurocardia lilliei ( c . a . fleming , 1943 ) † ( original combination )

lamarck j . b . ( 1801 ) . système des animaux sans vertèbres , ou tableau général des classes , des ordres et des genres de ces animaux ; présentant leurs caractères essentiels et leur distribution , d ' apres la considération de leurs rapports naturels et de leur organisation , et suivant l ' arrangement établi dans les galeries du muséum d ' histoire naturelle , parmi leurs dépouilles conservées ; précédé du discours d ' ouverture du cours de zoologie , donné dans le muséum national d ' histoire naturelle l ' an 8 de la république . published by the author and deterville , paris : viii + 432 pp . , available online at urltoken page ( s ) : 123 [ details ]

schmidt f . c . 1818 , versuch über die beste einrichtung zur aufstellung : behandlung und aufbewahrung der verschiedenen naturkörper und gegenṣtände der kunst , vorzüglich der conchylien - sammlungen , nebst kurzer beurtheilung der conchyliologischen systeme und schriften und einer tabellarischen zusammenstellung und vergleichung der sechs besten und neuesten conchyliologischen systeme . gotha , j . perth , 252 pp . , available online at urltoken ; seq = 15 page ( s ) : 57 [ details ]

( the eocene epoch began 57 . 8 million years ago and ended 36 . 6 million years ago ) . the shell , composed of two halves ( valves ) , is distinctive in form and generally large . transverse ribs radiate from the apex of the valves and are broken by a series of concentric growth rings . internally , the valves are marked by distinctive nodes along the edge and thickenings that form raised bars at the apex ; these form the surfaces along which the valves

eocene epoch , second of three major worldwide divisions of the paleogene period ( 66 million to 23 million years ago ) that began 56 million years ago and ended 33 . 9 million years ago . it follows the paleocene epoch and precedes the oligocene epoch . the eocene is often divided into early ( 56 million…

bivalve , ( class bivalvia ) , any of more than 15 , 000 species of clams , oysters , mussels , scallops , and other members of the phylum mollusca characterized by a shell that is divided from front to back into left and right valves . the valves are connected to one another at a hinge . primitive bivalves…

mollusk , any soft - bodied invertebrate of the phylum mollusca , usually wholly or partly enclosed in a calcium carbonate shell secreted by a soft mantle covering the body . along with the insects and vertebrates , it is one of the most diverse groups in the animal kingdom , with nearly 100 , 000 ( possibly…

gryphaea , extinct molluskan genus found as fossils in rocks from the jurassic period to the eocene epoch ( between 199 . 6 million and 33 . 9 million years ago ) . related to the oysters , gryphaea is characterized by its distinctively convoluted shape . the left valve , or shell , was much larger and more…

inoceramus , genus of extinct pelecypods ( clams ) found as fossils in jurassic to cretaceous rocks ( laid down between 199 . 6 million and 65 . 5 million years ago ) . especially important and widespread in cretaceous rocks , inoceramus had a distinctive shell ; it is large , thick , and wrinkled in a…

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horse , ( equus caballus ) , a hoofed , herbivorous mammal of the family equidae . it comprises a single species , …

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huber , m . ( 2010 ) . compendium of bivalves . a full - color guide to 3 , 300 of the world’s marine bivalves . a status on bivalvia after 250 years of research . hackenheim : conchbooks . 901 pp . , 1 cd - rom . ( look up in imis ) [ details ]

gofas , s . ; le renard , j . ; bouchet , p . ( 2001 ) . mollusca . in : costello , m . j . et al . ( eds ) , european register of marine species : a check - list of the marine species in europe and a bibliography of guides to their identification . patrimoines naturels . 50 : 180 - 213 . ( look up in imis ) [ details ]

zamouri - langar , n . ; chouba , l . ; ajjabi chebil , l . ; mrabet , r . ; el abed , a . ( 2011 ) . les coquillages bivalves des côtes tunisiennes . institut national des sciences et technologies de la mer : salammbô . isbn 978 - 9938 - 9512 - 0 - 2 . 128 pp . ( look up in imis ) [ details ]

jennifer hammock split the classifications by obis depth range resource , obis environmental information , and inventaire national du patrimoine naturel from cardites antiquatus ( linnaeus , 1758 ) to their own page .

hans - martin braun added the german common name " gefurchte trapezmuschel " to " cardites antiquatus ( linnaeus , 1758 ) " .

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see also camacho et al . 2000 , chiesa et al . 1995 , dockery 1982 , glenn 1904 , gliozzi and malatesta 1983 , heaslip 1968 , iqbal 1973 , lamprell and whitehead 1992 , le renard 1993 , marwick 1960 , olsson 1931 , sepkoski 2002 , vokes 1939 , vokes 1980 , vredenburg 1928 and woodring 1982

see also camacho et al . 2000 , gardner and bowles 1939 , glibert 1985 , olsson and richards 1961 , sepkoski 2002 , uliana and camacho 1975 , vokes 1939 and vokes 1980

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disclaimer : itis taxonomy is based on the latest scientific consensus available , and is provided as a general reference source for interested parties . however , it is not a legal authority for statutory or regulatory purposes . while every effort has been made to provide the most reliable and up - to - date information available , ultimate legal requirements with respect to species are contained in provisions of treaties to which the united states is a party , wildlife statutes , regulations , and any applicable notices that have been published in the federal register . for further information on u . s . legal requirements with respect to protected taxa , please contact the u . s . fish and wildlife service .

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calappoidea is a superfamily of crabs comprising the two families calappidae and matutidae .

the earliest fossils attributable to the calappoidea date from the aptian .

calappoidea

calappoidea <EOT> worms ( 2018 ) . calappoidea de haan , 1833 . accessed at : urltoken ; = 106694 on 2018 - 07 - 09

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1 : humboldt - universität zu berlin , institut für biologie / vergleichende zoologie , philippstr . 13 , d - 10115 berlin , germany ; technische universität ilmenau , institut für physik , weimarer str . 32 , d - 98684 ilmenau , germany

h . h . hobbs ; joan p . jass and jay v . huner

türkay , m . ( 2001 ) . decapoda , in : costello , m . j . et al . ( ed . ) ( 2001 ) . european register of marine species : a check - list of the marine species in europe and a bibliography of guides to their identification . collection patrimoines naturels , 50 : pp . 284 - 292 ( look up in imis ) [ details ]

martin , j . w . , & davis , g . e . ( 2001 ) . an updated classification of the recent crustacea . science series , 39 . natural history museum of los angeles county . los angeles , ca ( usa ) . 124 pp . ( look up in imis ) [ details ] available for editors [ request ]

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to cite this page : myers , p . , r . espinosa , c . s . parr , t . jones , g . s . hammond , and t . a . dewey . 2018 . the animal diversity web ( online ) . accessed at https : / / animaldiversity . org .

disclaimer : the animal diversity web is an educational resource written largely by and for college students . adw doesn ' t cover all species in the world , nor does it include all the latest scientific information about organisms we describe . though we edit our accounts for accuracy , we cannot guarantee all information in those accounts . while adw staff and contributors provide references to books and websites that we believe are reputable , we cannot necessarily endorse the contents of references beyond our control .

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lörenthey , e . & beurlen , k . , 1929 . die fossilen decapoden der länder der ungarischen krone . geologica hungarica , ( palaeontologica ) 3 : 1 - 421 , 12 tables , 16 pls .

linnaeus , c . v . , 1758 , systema naturae per regna tria naturae , secundum classes , ordines , genera , species , cum characteribus , differentiis , synonymis , locis ( ed . 10 ) , 1 : 1 - 824 . ( laurentii salvii , holmiae [ = stockholm ] ) .

calappidae - crsytal river fm . ( eocene ) - gainsville north florida usa .

ross , a . & scolaro , r . j . , 1964 . , a new crab from the eocene of florida . quarterly journal of the florida academy of sciences , 27 : 97 - 106 .

calappidae - priabonian ( upper eocene ) - parona , vicenza region , italy .

de angeli , a . & capriondo , fabio , 2009 . , crostacei decapodi del priaboniano di sossano ( monti berici , vicenza - italia settentrionale ) . studi e ricerche - associazione amici del museo - museo civico " g . zanatao " montecchio maggiore ( vicenza ) , 16 ( 2009 ) , pp . 23 - 33 .

calappidae - lutetian ( middle eocene ) - chiampo valley , vicenza , italy .

bittner , a . , 1886 , . neue brachyuren des eozäns von verona . sitzungsberichte der kaiserlichen akademie derwissenschaften inwien , ( mathematisch - naturwissenschaftliche klasse ) 94 ( 1 ) : 44 - 55 .

calappidae - castle hayne fm . ( eocene ) - pender county , north carolina , usa .

blow w . c . & . manning , r . c . , 1996 . preliminary descriptions of 25 new decapod crustaceans from the middle eocene of the carolinas , u . s . a . tulane studies in geology and paleontology , 29 ( 1 ) : 1 - 26 , pls . 1 - 5 .

calappidae - lutetian ( middle eocene ) - chiampo valley , vicenza , region , italy .

beschin , c . , busulini , a . , de angeli , a . , tessier , g . , 2002 , aggiornamento ai crostacei di cava “main” di arzignano ( vicenza – italia settentrionale ) ( crustacea , decapoda ) . studi e ricerche . studi e richerche , pp . 7 - 28 ,

calappidae - ? pleistocenes , ? holocene - okitsu , shimizu , suruga bay , shizuola pr . , japan .

de haan , w . , 1833 - 1850 . , crustacea . in : p . f . von siebold ( ed . ) , fauna japonica sive descriptio animalium , quae in itinere per japoniam , jussu et auspiciis superiorum , qui summum in india batava imperium tenent , suscepto , annis 1823 - 1830 collegit , notis , observationibus et adumbrationibus illustravit : i - xvii , i - xxxi , ix - xvi , 1 - 243 , pls . a - j , l - q , 1 - 55 , circ . table 2 . ( j . müller et co . , lugduni batavorum [ = leyden ] ) .

rathbun , m . j . , 1926 . , the fossil stalk - eyed crustacea of the pacific slope of north america . united states national museum bulletin , 138 : i - viii , 1 - 155 .

via , l . , 1959 . , decápodos fósiles del eoceno español . boletín del instituto geológico y minero de españa , 70 : 1 - 72 .

artal , p . & onetti , a . , 2017 . tavernolesia , new genus ( crustacea , decapoda , brachyura ) , from the eocene of the iberian peninsula . batalleria , 24 , 6 - 12 .

? - ilerdian ( lower ypresian [ lower eocene ] ) - isabena zone , huesca , aragon .

schweitzer , carrie e . & feldmann , rodney new species of calappid crabs from western north america and reconsideration of the calappidae sensu lato . journal of paleontology : vol . 74 , no . 2 , pp . 230 - 246 .

? aethridae - santee lms . fm . , eocene - south carloina , usa

maury , c . j . , , 1930 , o cretaceo da parahyba do norte . ministerio da agricultura , industria e commercio , serviço geologico e mineralogico do brasil , monographia , 8 : 1 - 305 .

? aethridae - lutetian ( middle eocene ) - chiampo valley , vicenza region , italy .

beschin , c . , busulini , a . , de angeli , a . & tessier , g . , 1994 , i crostacei eoceninci della cava " boschetto " di nogarole vicenttino ( vicenza - italia settentrionale ) . lavori - soc . ven . sc . nat - vol . 19 , pp . 159 - 215 , venezia , 31 gennaio 1994

bittner a . , 1875 , die brachyuren des vicentinischen tertiärgerbirges . denkschr . k . akad . wiss . , wien , 34 : 63 - 106 .

? aethridae - collbàs fm . , bartonian ( upper eocene ) - conca d ' òdena , anoia , catalonia .

de angeli , a . , beschin , c . , 1999 . i crostacei matutinae ( brachyura , calappidae ) delleocene del veneto ( italia settentrionale ) . studi e ricerche , pp 11 - 22 1999

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parent taxon : heterotremata according to c . e . schweitzer et al . 2010

see also fraaye and collins 1987 , tiwari et al . 1997 and todd and collins 2005

database contains : 10 . 643 species ( 763 with photo ) , 1 . 682 genera , 124 families

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chaloklum diving , 25 / 29 - 30 moo 7 , chaloklum village , koh phangan , suratthani 84280 . thailand .

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urn : lsid : biodiversity . org . au : afd . taxon : 85fcc08a - 0539 - 44be - 9692 - cf273716f8a7

urn : lsid : biodiversity . org . au : afd . taxon : 0654b9d2 - e4b4 - 4e49 - ae73 - 520d8c1d839f

urn : lsid : biodiversity . org . au : afd . name : 547390

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