the pale chub , ( zacco platypus ) , also known as pale bleak or fresh-water sprat , is a species of fish native to rivers and streams from northern china and korea to northern vietnam .

they can grow up to 20 centimetres ( 7.9 in ) but usually grow up to 13 centimetres ( 5.1 in ) .

its diet consists of zooplankton , invertebrates , fish , and debris .

zacco platypus is called oikawa オイカワ （ 追河 、 opsariichthys platypus ） in japan .

zacco platypus

zacco platypus <EOT> huangshan population of chinese zacco platypus ( teleostei , cyprinidae ) harbors diverse matrilines and high genetic diversity .

japanische drachenfische ( zacco platypus ) laichen im gartenteich unseres kunden benedikt reisner ab . weitere informationen unter : urltoken

sympatric fish species include sicyopterus japonicus , spinibarbus hollandi , acrossocheilus paradoxus , varicorhinus barbatulus , varicorhinus alticorpus , zacco pachycephalus , zacco platypus , aphyocypris kikuchii and hemimyzon taitungensis .

huangshan population of chinese zacco platypus ( teleostei , cyprinidae ) harbors diverse matrilines and high genetic diversity . - pubmed - ncbi

intron polymorphism ( epic - pcr ) reveals phylogeographic structure of zacco platypus in china : a possible target for aquaculture development .

aggressive interactions between the dark chub , zacco temmincki , and the pale chub , z . platypus , in relation to their feeding behaviour

intron polymorphism ( epic - pcr ) reveals phylogeographic structure of zacco platypus in china : a possible target for aquaculture development . - pubmed - ncbi

" aggressive interactions between the dark chub , zacco temmincki , and the pale chub , z . platypus , in relation to their feeding behaviour "

it is important to make clear seasonal changes of fish habitat quality in rivers . there are still many unsolved question in fish habitat quality , because this consists of quite complex system . in this study , field surveys of seasonal changes of fish habitat quality were conducted through nine months in itabitsugawa river which belongs to class b river . it was found that carassius langsdorfi moves from pool to riffle through winter to summer . zacco temminckii and zacco platypus evade carassius langsdorfi in all seasons , because the body length of carassius langsdorfi is larger than that of zacco temminckii and zacco platypus . further , zacco temminckii evades zacco platypus which is larger than 8cm of body length .

aggressive interactions between the dark chub , zacco temmincki , and the pale chub , z . platypus , in relation to their feeding behaviour [ 1994 ]

the floods sometimes occur so that the discharge , velocity , flow depth are changed . in such situations , fish behaviors may be changed . however , there is little information on the fish behaviors in flood . in this study , the swimming behaviors of zacco platypus ( oikawa ) in the increasing discharge flows were recorded with a digital video camera . it was found that swimming speed of zacco platypus increased with an increase of velocity . in contrast , the ground speed of zacco platypus did not so change , irrespective of increasing discharge . the ground speed and swimming distance of zacco platypus did not change , even if the acceleration of velocity is changed . the effects of the acceleration on the swimming behavior of zacco platypus was smaller than that of the velocity .

phylogenetic structure of zacco platypus ( teleostei , cyprinidae ) populations on the upper and middle chang jiang ( = yangtze ) drainage inferred from cytochrome b sequences .

reproduction of the oikawa , zacco platypus ( temminck and schlegel ) , a cyprinid i . sexual characters in the anal fin and maturation kenya mizuguchi and yoshio hiyama

phylogenetic structure of zacco platypus ( teleostei , cyprinidae ) populations on the upper and middle chang jiang ( = yangtze ) drainage inferred from . . . - pubmed - ncbi

matrix of pairwise genetic variation between matrilines ( a - j ) of z . platypus

influence of glaciation on divergence patterns of the endemic minnow , zacco pachycephalus , in taiwan .

berrebi p et al . intron polymorphism ( epic - pcr ) reveals phylogeographic structure of zacco platypus in china : a possible target for aquaculture development . heredity ( edinb ) . 2005 ; 94 : 589 - 98

spawning characteristics between the two populations were not observed . bases on the biology characteristics of zacco platypus in beijing , we suggests to control the fishing amount , choose comfortable fishing season and protect living condition in order to maintain

it is important to secure the rest area for fish . fish has the ordinary and dark muscle . when fish uses ordinary muscle , fish gets tired . in such a situation , fish needs a rest . in this study , vegetation density in open - channel is changed . the trajectories of zacco platypus ' s were observed . it was found that zacco platypus utilized a slow of the speed as rest space . stay in vegetation area increases with an increase of vegetation density . an approach rate to vegetation area rises with an increase of vegetation density . however , it was not found that clear relationship with a migration rate of zacco platypus and vegetation density .

six main mitochondrial dna ( mtdna ) lineages have been described in minnow ( zacco platypus ) samples obtained from northern , western and southern china . perdices et al . ( 2004 ) predicted that further sampling of other tributaries might discover more lineages of this species . in this study , we collected 26 zacco platypus individuals in the huangshan area of eastern china and determined the cytochrome b ( cytb ) sequence variations . combined with reported data in genbank , we identified ten matrilines ( zacco a - j ) in a total of 169 samples , with relatively high molecular divergence found among them . the huangshan population had the greatest genetic variation among all sampled regions and hosted six of the ten matrilines . our results highlight the significance of the huangshan area for the conservation of zacco platypus .

berrebi p & boissin e & fang f , et al . ( 2005 ) . intron polymorphism ( epic - pcr ) reveals phylogeographic structure of zacco platypus in china : a possible target for aquaculture development . heredity , 94 , pp . 589 - 98 .

berrebi p et al : " intron polymorphism ( epic - pcr ) reveals phylogeographic structure of zacco platypus in china : a possible target for aquaculture development . " heredity , vol . 94 , no . 6 , 2005 , pp . 589 - 98 , urltoken accessed july 9 , 2018 .

berrebi p , boissin e , fang f , et al . intron polymorphism ( epic - pcr ) reveals phylogeographic structure of zacco platypus in china : a possible target for aquaculture development . heredity ( edinb ) 2005 ; 94 ( 6 ) : 589 - 98 urltoken accessed july 9 , 2018 .

we examined the genetic structure and phylogenetic relationships of some chinese populations from the chang jiang ( = yangtze ) drainage of the cyprinid zacco platypus . we sequenced the complete mitochondrial cytochrome b gene of 64 individuals from 6 upper and middle tributaries of the sichuan and hunan provinces to assess their population structure and systematics . the combined analyses of the phylogenetic information and the population structure suggested that chinese z . platypus consist of four distinct mtdna lineages which exhibit high genetic variation and haplotypic diversity ( zacco a - d ) . the high molecular divergence observed among zacco a - d mtdna lineages ( trn + i ( 0 . 76 ) distance , mean 8 . 9 % + / - 1 . 7 % ) and their phylogeographic structure indicate that all four lineages have evolved independently . analysis of molecular variance ( amova ) indicates that most of the genetic variation observed is found among the four zacco mtdna lineages ( thetact = 0 . 94 ) suggesting restricted gene flow among the chang jiang populations . long - term interruption of gene flow was also evidenced by thetast values higher than 0 . 9 that could be favoured by the discontinuous distributions of the lineages inhabiting upper ( sichuan province ) and middle ( hunan province ) chang jiang tributaries . the significant correlation between the geographic and genetic distances provide support for the importance of geographic discontinuity in shaping the zacco genetic structure . nested clade analysis ( nca ) results were congruent with phylogenetic relationships recovered and confirm the genetic distinctiveness of four independent zacco groups . these four groups correspond to the four zacco a - d mtdna lineages recovered in the phylogeny and were defined by nucleotide synapomorphies permitting bootstrapped and bayesian confidence of 95 % or greater . the high level of mitochondrial sequence divergence separating all zacco mtdna lineages suggested that the z . platypus populations from the chang jiang drainage probably correspond to four different species .

perdices et al . ( 2004 ) and perdices & coelho ( 2006 ) divided z . platypus sampled in southern , western and northern china into matrilines a–f . they suggested that the long - term interruption of gene flow might have caused the diversification and an underestimation of the number of species . our analyses identified ten matrilines of z . platypus in chinese and some japanese populations . this confirms the prediction of perdices et al . ( 2004 , page : 9 ) that “exhaustive sampling of other tributaries might evidence other zacco lineages” . this is also in accordance with that found for opsariichthys bidens , a sympatric species of z . platypus ( perdices et al . , 2005 ) .

zacco platypus , pale chub , is an indigenous freshwater fish of east asia including korea and has many useful characteristics as indicator species for water pollution . while utility of z . platypus as an experimental species has been recognized , genetic - level information is very limited and warrants extensive research . metallothionein ( mt ) is widely used and well - known biomarker for heavy metal exposure in many experimental species . in the present study , we cloned mt in z . platypus and evaluated its utility as a biomarker for metal exposure . for this purpose , we sequenced complete complementary dna ( cdna ) of mt in z . platypus and carried out phylogenetic analysis with its sequences . the transcription - level responses of mt gene following the exposure to cdcl 2 were also assessed to validate the utility of this gene as an exposure biomarker . analysis of cdna sequence of mt gene demonstrated high conformity with those of other fish . mt messenger rna ( mrna ) expression and enzymatic mt content significantly increased following cdcl 2 exposure in a concentration - dependent manner . the level of cdcl 2 that resulted in significant mt changes in z . platypus was within the range that was reported from other fish . the mt gene of z . platypus sequenced in the present study can be used as a useful biomarker for heavy metal exposure in the aquatic environment of korea and other countries where this freshwater fish species represents the ecosystem .

zacco platypus is a common minnow that occurs in sympatry with most chinese cyprinids ( deng et al . , 2013 ) . topographical barriers may restrict its life history and drive cryptic diversity . the species’ distribution encompasses all major river systems in mainl and china , as well as the korean peninsula and japan ( chen , 1998 ) . perdices et al . ( 2004 ) analyzed the genetic diversity of z . platypus sampled in the upper and middle changjiang ( yangtze river ) and found four major matrilines that may harbor multiple species . long - term interruption of dispersal is thought to have driven this diversity . perdices and coelho ( 2006 ) further studied samples from the pearl river and northern drainages , and obtained six matrilines in china . using nuclear dna data , berrebi et al . ( 2005 ) identified four genetic groups within z . platypus from sichuan , hunan and guangxi provinces in china .

some drainages still await sampling , such as the yellow river , one of the most important drainages in china . future research should detect additional matrilines of zacco , while morphological analyses may help differentiate morphological differences of taxonomic significance .

interpreted four basic population history scenarios based on haplotype and nucleotide diversities , which can also be used to clarify the history of z . platypus populations . our results revealed a pattern of high haplotype and nucleotide diversity in the huangshan population (

ty - jour t1 - intron polymorphism ( epic - pcr ) reveals phylogeographic structure of zacco platypus in china : a possible target for aquaculture development . au - berrebi , p , au - boissin , e , au - fang , f , au - cattaneo - berrebi , g , py - 2005 / 6 / 9 / pubmed py - 2005 / 9 / 15 / medline py - 2005 / 6 / 9 / entrez sp - 589 ep - 98 jf - heredity jo - heredity ( edinb ) vl - 94 is - 6 n2 - the present study aims at a phylogeographic description of zacco platypus from southeast china , in order to detect subdivisions within the nominal species . two main basins were sampled : the chang jiang ( yangstze river ) in central and east china ( hunan and sichuan provinces ) and the xi jiang , the more southern main tributary of the zhu jiang ( pearl river , guangxi province ) . a total of 27 intron systems were tested , five of them were informative and gave 12 interpretable and polymorphic loci . within the diversity of z . platypus , four genetic groups were identified by multidimensional ( fca ) analyses , corresponding to distinct genetic pools . the geographical distribution of the genetic groups corresponds neither with the drainage structure , nor the geographic distances between samples . it follows that isolation by distance and limited migration are insufficient to explain this geographic structure . the history of the river network therefore appears to have played an important role . sn - 0018 - 067x ur - urltoken l2 - urltoken er -

) . the haplotypes were grouped into main clade 1 and 2 . clade 1 hosted individuals from huangshan , sichuan , hunan and guangxi and clade 2 contained specimens from huangshan , beijing and japan . we identified ten matrilines of z . platypus according to the topology of the phylogenetic tree and the genetic variation between the ten matrilines .

we used 916 bp out of 1 140 bp of the cytb sequences in the following analyses . the newly obtained sequences and those downloaded from genbank were aligned using clustal x ( thompson et al . , 1997 ) . for phylogenetic reconstruction , two closely related species , zacco temmincki and c and idia barbatus ( mayden et al . , 2007 ; wang et al . , 2011 ) , were chosen as outgroups .

in view of modern genetics , genetic diversity in a given species is closely related to its adaptability , variability , and evolutionary potentiality , with genetic variation considered a prerequisite for organisms to cope with environmental uncertainty ( conrad , 1983 ) . herein , we report on the genetic diversity of z . platypus from eastern china based on extensive sampling of the huangshan area together with prior cytb sequence data from mainl and china , taiwan ( perdices et al . , 2004 ; perdices & coelho , 2006 ; wang et al . , 2007 ) , and japan ( he et al . , 2004 ; kawamura et al . , 2014 ; kitamura et al . , 2012 ; sasaki et al . , 2007 ; wang et al . , 2007 ) . we further evaluated the matrilineal diversity of z . platypus and revealed the possible ecological significance of the huangshan area .

reproduction and organ structure of the pale chub , zacco platypus in isa stream were investigated by means of histological methods . the results of the study confirmed reproductive abnormality and histopathological features in the pale chub . the gonadosomatic index ( gsi ) of the fish showed two peak in april and august . in summer season , gsi of the male was about two times of the female’s gsi . monthly variation of the gonadal development was very irregular . from the histological analysis of the organ structure , epidermal atrophy , necrosis and hyperplasia of pigment cell were observed in the skin . epithelial layer lifting and clubbing of the lamella and bifurcation of the filament were observed in the gill . also histological changes as congestion , cytoplasmic degeneration of hepatic cell , degeneration of bile duct , glomerular dilatation , degeneration of renal tubule and pycnosis of interstitial cell were identified in the liver and kidney , respectively .

values above branches represent the support level of ml ( bsp ) and values below branches represent the popularity rating of bi ( bpp ) . vertical bars indicate the mtdna lineage assignment ( a - j ) . < italic > zacco < / italic > a - f follow the nomenclature of < xref ref - type = " bibr " rid = " b23 - zoolres - 37 - 2 - 103 " > perdices & coelho ( 2006 ) < / xref > and the others follow the alphabet . bold types are huangshan populations .

the oikawa , zacco platypus , is the popular game fish which widely inhabits the rivers and ponds all through japan except hokkaido . along with the study on the life - history , ecology and population of the species , which has been carried out since 1965 mainly in the water of the river aki ( a tributary of the river tama flowing into tokyo bay ) , the authors have studied the relationships between the maturity and sexual characters in the anal fin . based on the inspection of 1628 specimens collected monthly by a cast - net from november 1965 to september 1966 and in may 1967 , and by angling in august 1966 , the following results were obtained . 1 ) the males during the spawning season ( june to august ) were characterized by well - developed pearl organs , red and bluish green color on the side and enormously grown anal fin . these mature males , dominated by age group iii + and a half of males of age group ii + , exhibited breeding reactions and died out no later than september after spawning . 2 ) on the other hand , another half of males of age group ii + continued to grow in summer and became matured at iii + year old in the following summer . these ii + males were termed “non - mature” males against the above mentioned mature males . these males showed no secondary sexual characteristics and having only thin thread - like testis through out the spawning season . 3 ) the morphological change of anal fin was observed also in mature females , but to a lesser extent as compared with mature males . mature females began to spawn at ii + years and died out after the second spawning at iii + years . 4 ) it is evident that zacco platypus more than 80 mm in total length can be sexed during spawning season and the degree of the sexual maturation , particularly in the males , may be quantitatively measured by the length of anal fin . 5 ) the cause of the condition that males of age group ii + are mature or “non - mature” in spawning season was suggested through reading the history on the scales examined . the appearance of “non - mature” males in age group ii + was discussed in relation to the population density .

the effects of effluent from a municipal wastewater treatment plant ( mwtp ) on population level responses of the pale chub ( zacco platypus ) , as sentinel species , were evaluated at four sites of the gap stream ( gs ) in june 2007 . at gs 7 . 2 and gs 2 . 7 , downstream of the municipal wastewater outfall , the response patterns of the pale chub population in the age at maturation were not changed ; mean age , fecundity , and condition factor were increased ; age distribution was shifted to older ; growth rate was increased / or not changed ; and the egg size and population size were decreased compared with reference site ( gs 26 . 6 ) . these observed responses of the pale chub population matched well with the characteristics expected from a fish population experiencing chronic recruitment failure , except for the decreased egg size due to the elevation in nutrients and pollutants by treated sewage discharge . the observed response pattern at the downstream sites might be caused by the deterioration of the spawning or nursery habitat , or by the contaminant - induced chronic spawning failures . thus , these results suggest that the effluent of the mwtp might impact on the fish population structure and health status of pale chub population .

divergence times among the main lineages of z . platypus were estimated using a bayesian mcmc approach implemented in beast v . 1 . 7 . 5 based on a strict molecular clock ( drummond & rambaut , 2007 ) . the parameters were : substitution model , tn93 + g ; tree prior , coalescent : constant size ; normal distribution ; 10 million generations ; parameters logged every 1 000 ; burn - in value = 1 000 . the molecular clock of cyprinids was assumed to be 1 . 52 % site - 1 ma ( million years ) - 1 ( doadrio et al . , 2002 ) for cyt b .

theory indicates that the anti - predation behavior of a fish species will depend on predation threats in a way that tends to improve the probability of surviving predation . pale chub ( zacco platypus ) , a small asiatic cyprinid , is widely distributed in both high - and low - predation reaches of the wujiang river . to test whether the anti - predator behavior of pale chub along the wujiang river varied in fish living in habitats under different levels of predation pressure , we measured spontaneous swimming activity , risk - taking behavior , utilization of shelter and boldness in pale chub collected from both high - and low - predation habitats in the wujiang river . the fish from high - predation populations showed less spontaneous swimming activity and risk - taking behavior compared with those of the fish from low - predation populations . however , neither utilization of shelter nor boldness exhibited any significant differences between high - and low - predation populations . one reason for this result may be that in their daily lives , the pale chub in the high - predation population primarily respond to the presence of predators by decreasing spontaneous swimming activity and risk - taking behaviors when threatened rather than increasing utilization of shelters and showing less bold behavior . the lack of an increase in shelter utilization and the lack of a decrease in boldness in response to predation pressure may be a compromise to allow foraging and growth .

fish inhabit environments that vary greatly in terms of predation intensity , and these predation regimes are generally expected to be a major driver of divergent natural selection . to test whether there is predator - driven intra - species variation in the locomotion , metabolism and water velocity preference of pale chub ( zacco platypus ) along a river , we measured unsteady and steady swimming and water velocity preference among fish collected from both high - and low - predation habitats in the wujiang river . we also measured the routine metabolic rate ( rmr ) , maximum metabolic rate ( mmr ) and cost of transport ( cot ) and calculated the optimal swimming speed ( u opt ) . the fish from the high - predation populations showed a shorter response latency , elevated routine metabolism , lower swimming efficiency at low swimming speed , and lower water velocity preference compared with those from the low - predation populations . neither of the kinematic parameters fast - start and critical swimming speed ( u crit ) showed a significant difference between the high - and low - predation populations . the fish from the high - predation populations may improve their predator avoidance capacity primarily through an elevated routine metabolism and shorter response latency to achieve advanced warning and escape , rather than an improved fast - start swimming speed or acceleration . thus , the cost of this strategy is an elevated rmr , and no trade - off between unsteady and steady swimming performance was observed in the pale chub population under various predation stresses . it was interesting to find that the high - predation fish showed an unexpected lower velocity preference , which might represent a compromise between predation avoidance , foraging and energy saving .

although perdices et al . ( 2004 ) predicted that exhaustive sampling of other tributaries might discover other lineages of zacco , few specimens have been sampled in eastern china . the huangshan area in eastern china is a mosaic of mountains with elevations lower than 2 000 m , and exhibits a complex geological history that includes tectonic movements , orogenesis , and periodic climatic change ( e . g . , ju et al . , 2007 ; rüber et al . , 2004 ; zhang et al . , 1990 ) . based on patterns of intraspecific genetic variation and buffer - zone models , huangshan hosts refugia of eastern asian conifers , frogs , non - migratory birds and asian salam and ers ( gao et al . , 2007 ; li et al . , 2009 ; murphy et al . , 2000 ; wu et al . , 2013 ; zhang et al . , 2008 ) .

maturity : l m ? range ? - ? cm max length : 20 . 0 cm tl male / unsexed ; ( ref . 5258 ) ; common length : 13 . 0 cm sl male / unsexed ; ( ref . 35840 )

adults are common in streams and rivers with rapid water flow , but not deep or stagnant waters . feed on zooplankton , small crustaceans , macroscopic algae , small fish and detritus .

man , s . h . and i . j . hodgkiss , 1981 . hong kong freshwater fishes . urban council , wishing printing company , hong kong , 75 p . ( ref . 5258 )

phylogenetic diversity index ( ref . 82805 ) : pd 50 = 0 . 5625 [ uniqueness , from 0 . 5 = low to 2 . 0 = high ] .

bayesian length - weight : a = 0 . 00741 ( 0 . 00575 - 0 . 00955 ) , b = 3 . 12 ( 3 . 05 - 3 . 19 ) , in cm total length , based on lwr estimates for this species ( ref . 93245 ) .

trophic level ( ref . 69278 ) : 3 . 1 ±0 . 43 se ; based on food items .

resilience ( ref . 69278 ) : medium , minimum population doubling time 1 . 4 - 4 . 4 years ( assuming fec < 10 , 000 ) .

vulnerability ( ref . 59153 ) : low to moderate vulnerability ( 30 of 100 ) .

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head moderately large . eye moderately large , on upper side of head . mouth oblique ; posterior end of upper jaw reaches anterior margin of eye . body elongated , posterior laterally compressed ; dorsal and ventral profile slightly arched . barbels absent . scales moderately large , cycloid ; lateral line complete and curved above pelvic fin origin ; head naked . l . l . ：43 - 45 ; predorsal scales：16 - 18 ; dorsal fin rays：3 + 7 ; pelvic fin rays：1 + 8 ; anal fin rays：3 + 8 - 9 . dorsal fin origin near midline between snout tip and caudal peduncle end ; pectoral fin end distant from pelvic fin origin ; pelvic fin at lower side of body ; anal fin rays long , the longest reaches caudal peduncle end ; caudal fin forked . dark grayish dorsally , side and belly silvery white . 10 or more vertically bluish stripes on side of spawning male .

primary freshwater fish , prefers running water at middle reaches . occasionally form schools . polyphagia , feeds on small aquatic animals and diatom .

extensively distributed to eastern asia , including eastern china , korea , japan and taiwan . populations of taiwan were restricted in northwestern taiwan .

bayesian length - weight : a = 0 . 00741 ( 0 . 00575 - 0 . 00955 ) , b = 3 . 12 ( 3 . 05 - 3 . 19 ) , in cm total length , based on lwr estimates for this species ( ref .

) : 3 . 1 ±0 . 43 se ; based on food items .

) : medium , minimum population doubling time 1 . 4 - 4 . 4 years ( assuming fec < 10 , 000 ) .

html public " - / / w3c / / dtd xhtml 1 . 0 strict / / en " " urltoken "

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special publication of the center for biodiversity research and information , no . 1 , vol 1 - 3

disclaimer : itis taxonomy is based on the latest scientific consensus available , and is provided as a general reference source for interested parties . however , it is not a legal authority for statutory or regulatory purposes . while every effort has been made to provide the most reliable and up - to - date information available , ultimate legal requirements with respect to species are contained in provisions of treaties to which the united states is a party , wildlife statutes , regulations , and any applicable notices that have been published in the federal register . for further information on u . s . legal requirements with respect to protected taxa , please contact the u . s . fish and wildlife service .

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centro de biologia ambiental , departamento de zoologia e antropologia , faculdade de ciências , bloco c - 2 , 3 piso , campo grande , 1749 - 016 lisbon , portugal . anabelperdices @ urltoken

zheng x 1 , zhou tq 1 , wan t 2 , perdices a 3 , yang jq 4 , tang xs 5 , wang zp 6 , huang lq 7 , huang s 8 , he sp 9 .

key laboratory of exploration and utilization of aquatic genetic resources , shanghai ocean university , ministry of education , shanghai 201306 , china ; college of life and environment sciences , huangshan university , huangshan anhui 245041 , china .

state key laboratory of genetic resources and evolution , kunming institute of zoology , chinese academy of sciences , kunming yunnan 650223 , china .

museo nacional de ciencias naturales , csic , c / josé gutiérrez abascal , 2 , 28006 madrid , spain .

key laboratory of exploration and utilization of aquatic genetic resources , shanghai ocean university , ministry of education , shanghai 201306 , china .

college of life and environment sciences , huangshan university , huangshan anhui 245041 , china .

college of life and environment sciences , huangshan university , huangshan anhui 245041 , china ; state key laboratory of genetic resources and evolution , kunming institute of zoology , chinese academy of sciences , kunming yunnan 650223 , china . snakeman @ hsu . edu . cn .

institute of hydrobiology , chinese academy of sciences , wuhan hubei 430072 , china . clad @ ihb . ac . cn .

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found in shallow fast - flowing streams and rivers in asia from northern china to vietnam .

this page was last edited on 13 december 2017 , at 03 : 09 .

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jennifer hammock added an association between " yangtze river benthopelagic habitat " and " saurogobio lissilabris banarescu & nalbant 1973 " .

jennifer hammock added an association between " yangtze river benthopelagic habitat " and " pareuchiloglanis sinensis ( hora & silas , 1952 ) " .

jennifer hammock added an association between " yangtze river benthopelagic habitat " and " glyptothorax sinensis ( regan , 1908 ) " .

jennifer hammock added an association between " yangtze river benthopelagic habitat " and " glyptothorax fokiensis ( rendahl , 1925 ) " .

jennifer hammock added an association between " yangtze river benthopelagic habitat " and " hucho bleekeri kimura , 1934 " .

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journal of japan society of civil engineers , ser . b1 ( hydraulic engineering )

edited and published by : japan society of civil engineers produced and listed by : iword co . , ltd .

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to cite this page : myers , p . , r . espinosa , c . s . parr , t . jones , g . s . hammond , and t . a . dewey . 2018 . the animal diversity web ( online ) . accessed at https : / / animaldiversity . org .

disclaimer : the animal diversity web is an educational resource written largely by and for college students . adw doesn ' t cover all species in the world , nor does it include all the latest scientific information about organisms we describe . though we edit our accounts for accuracy , we cannot guarantee all information in those accounts . while adw staff and contributors provide references to books and websites that we believe are reputable , we cannot necessarily endorse the contents of references beyond our control .

this material is based upon work supported by the national science foundation grants drl 0089283 , drl 0628151 , due 0633095 , drl 0918590 , and due 1122742 . additional support has come from the marisla foundation , um college of literature , science , and the arts , museum of zoology , and information and technology services .

20 . 0 cm tl ( male / unsexed ; ( ref . 5258 ) )

common in streams and rivers with rapid water flow , but not deep or stagnant waters . feeds on zooplankton , small crustaceans , macroscopic algae , small fish and detritus .

the following is a representative barcode sequence , the centroid of all available sequences for this species . there is 1 barcode sequence available from bold and genbank .

below is the sequence of the barcode region cytochrome oxidase subunit 1 ( coi or cox1 ) from a member of the species .

mitochondrial diversity of opsariichthys bidens ( teleostei , cyprinidae ) in three chinese drainages .

molecular phylogeography and population structure of a mid - elevation montane frog leptobrachium ailaonicum in a fragmented habitat of southwest china .

exploring factors shaping population genetic structure of the freshwater fish sinocyclocheilus grahami ( teleostei , cyprinidae ) .

chloroplast dna phylogeography of clintoniaudensis trautv . & mey . ( liliaceae ) in east asia .

phylogeography of the endemic gymnocypris chilianensis ( cyprinidae ) : sequential westward colonization followed by allopatric evolution in response to cyclical pleistocene glaciations on the tibetan plateau .

molecular phylogeography of endangered sharp - snouted pitviper ( deinagkistrodon acutus ; reptilia , viperidae ) in mainland china .

this study was supported by the center for aquatic ecosystem restoration ( caer ) of eco - star project from ministry of environment ( moe , korea ) . s . lee is supported by the national research foundation of korea .

amiard , j . c . , amiard - triquet , c . , barka , s . , pellerin , j . , & rainbow , p . s . ( 2006 ) . metallothioneins in aquatic invertebrates : their role in metal detoxification and their use as biomarkers .

bervoets , l . , knapen , d . , de jonge , m . , van campenhout , k . , & blust , r . ( 2013 ) . differential hepatic metal and metallothionein levels in three feral fish species along a metal pollution gradient .

coyle , p . , philcox , j . c . , carey , l . c . , & rofe , a . m . ( 2002 ) . metallothionein : the multipurpose protein .

de martinez gaspar martins , c . , & bianchini , a . ( 2009 ) . metallothionein - like proteins in the blue crab

eaton , d . l . , & toal , b . f . ( 1982 ) . evaluation of the cd / hemoglobin affinity assay for the rapid determination of metallothionein in biological tissues .

hall , t . a . ( 1999 ) . bioedit : a user - friendly biological sequence alignment editor and analysis program for windows 95 / 98 / nt .

haq , f . , mahoney , m . , & koropatnick , j . ( 2003 ) . signaling events for metallothionein induction .

( siluriformes ) metallothionein cdna : molecular cloning and transcript expression level in response to exposure to the heavy metals cd , cu , and zn .

lange , a . , ausseil , o . , & segner , h . ( 2002 ) . alterations of tissue glutathione levels and metallothionein mrna in rainbow trout during single and combined exposure to cadmium and zinc .

livak , k . j . , & schmittgen , t . d . ( 2001 ) . analysis of relative gene expression data using real - time quantitative pcr and the 2 ( - delta delta c ( t ) ) method .

mathiessen , p . ( 2000 ) . biological effects quality assurance in monitoring programs ( belqualm ) . in centre for environment faasc , ed . , remembrance avenue , burnham - on - crouch , essex smo 8ha , uk .

rodriguez - ortega , m . j . , alhama , j . , funes , v . , romero - ruiz , a . , rodriguez - ariza , a . , & lopez - barea , j . ( 2002 ) . biochemical biomarkers of pollution in the clam

ross , k . , cooper , n . , bidwell , j . r . , & elder , j . ( 2002 ) . genetic diversity and metal tolerance of two marine species : a comparison between populations from contaminated and reference sites .

tsuda , t . , inoue , t . , kojima , m . , & aoki , s . ( 1996 ) . pesticides in water and fish from rivers flowing into lake biwa .

tsuda , t . , kojima , m . , harada , h . , nakajima , a . , & aoki , s . ( 1998 ) . pesticides and their oxidation products in water and fish from rivers flowing into lake biwa .

yeom , d . h . , lee , s . a . , kang , g . s . , seo , j . , & lee , s . k . ( 2007 ) . stressor identification and health assessment of fish exposed to wastewater effluents in miho stream , south korea .

zrnčić , s . , oraić , d . , ćaleta , m . , mihaljević , z . , zanella , d . , & bilandžić , n . ( 2013 ) . biomonitoring of heavy metals in fish from danube river .

lee , s . , kim , c . , kim , j . et al . environ monit assess ( 2015 ) 187 : 447 . urltoken

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photo credit : t . - c . francis pan some oyster larvae grow faster than others , but now donal t . manahan and colleagues reveal that the fastest growers are marked out by their high protein synthesis rates .

“one of the underappreciated benefits of fellowships is the act of applying for them , because you have to write and articulate your ideas . ”

in our latest early - career interview , we chat to simon sponberg , assistant professor at the georgia institute of technology , usa . he shares the story of his career , beginning with how he combined a love of physics and biology by studying how geckos stick to walls .

a new review from craig p . mcgowan and clint e . collins looks at the ecology , biomechanics and evolution of bipedal hopping in mammals , with a focus on why bipedal hopping has arisen in multiple clades of mammals .

photo credit : ari friedlaender not all orca species prey on aquatic mammals , so how do delphinids know when they are at risk ? a new study shows that pilot whales and risso’s dolphins flee from a subset of orca calls that share acoustic characteristics with other mammal alarm calls , including human screams . this article was featured in science magazine .

at the heart of prelights is the community of early - career researchers who select and highlight interesting preprints in various fields . we are now ready to grow our team of prelighters who are passionate about preprints and enjoy writing and communicating science . find out more here and apply by the extended deadline , 20 july 2018 .

freshwater minnow is a primary freshwater fish usually found in clean streams and rivers . the fish has a long and compressed body . there is no barbel on its terminal mouth . its bosy is light to brownish yellow with 12 to 13 silvery vertical bars on its sides . ther male shows elongated rays in its anal fin and yellow peral organs on the jaws and opercula during breeding season

lam , k . s . ( 2002 ) . freshwater fish in hong kong . agriculture , fisheries and conservation department , friends of the country parks and cosmos books ltd . , hong kong .

lee , v . l . f . , lam , s . k . s . , ng , f . k . y . , chan , t . k . t . and young , m . l . c . ( 2004 ) . field guide to the freshwater fish of hong kong . agriculture , fisheries and conservation department , friends of the country parks and cosmos books ltd . , hong kong .

man , s . h . and hodgkiss , i . j . ( 1981 ) . hong kong freshwater fishes . the urban council , hong kong .

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vasil ' ev , v . p . , 1980 . chromosome numbers in fish - like vertebrates and fish . j . ichthyol . 20 ( 3 ) : 1 - 38 .

man , s . h . and i . j . hodgkiss , 1981 . hong kong freshwater fishes . urban council , wishing printing company , hong kong , 75 p .

hwang , h . c . , p . c . yueh and s . f . yu , 1982 . the freshwater fishes of china in colored illustrations . vol . 1 . shanghai sciences and technology press , shanghai , china . 173 p .

masuda , h . , k . amaoka , c . araga , t . uyeno and t . yoshino , 1984 . the fishes of the japanese archipelago . vol . 1 . tokai university press , tokyo , japan . 437 p . ( text ) .

takai , a . and y . ojima , 1984 . some features on the nucleolus organizer regions in the chromosomes of the cyprinid fishes . proc . japan acad . , ser . b : phys . biol . sci . 60 ( 10 ) : 410 - 413 .

li , y . , k . li , y . hong , j . gui and d . zhou , 1985 . studies on the karyotypes of chinese cyprinid fishes . vii . karyotypic analyses of seven species in the subfamily leuciscinae with a consideration for the phylogenetic relationships of some cyprinid fishes concerned . acta genet . sin . 12 ( 5 ) : 367 - 372 .

nagata , y . and y . nakata , 1988 . distribution of six species of bitterlings in a creek in fukuoka prefecture , japan . jap . j . ichthyol . 35 ( 3 ) : 320 - 331 .

hureau , j . - c . , 1991 . la base de données gicim : gestion informatisée des collections ichthyologiques du muséum . p . 225 - 227 . in atlas préliminaire des poissons d ' eau douce de france . conseil supérieur de la pêche , ministère de l ' environnement , cemagref et muséum national d ' histoire naturelle , paris .

shao , k . - t . and p . l . lim , 1991 . fishes of freshwater and estuary . encyclopedia of field guide in taiwan . recreation press , co . , ltd . , taipei . vol . 31 . 240 p . ( in chinese ) .

ye , f . and p . song , 1991 . danioninae . p . 67 - 79 . in j . - h . pan , l . zhong , c . - y . zheng , h . - l . wu and j . - h . liu ( eds ) . 1991 . the freshwater fishes of guangdong province . guangdong science and technology press , guangzhou . 589 pp .

shen , s . c . ( ed . ) , 1993 . fishes of taiwan . department of zoology , national taiwan university , taipei . 960 p .

okiyama , m . , 1993 . an atlas of the early stage fishes in japan . koeltz scientific books , germany . 1154 p .

baensch , h . a . and r . riehl , 1995 . aquarien atlas . band 4 . mergus verlag gmbh , verlag für natur - und heimtierkunde , melle , germany . 864 p .

klinkhardt , m , m . tesche and h . greven , 1995 . database of fish chromosomes . westarp wissenschaften .

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eschmeyer , w . n . ( ed . ) , 1998 . catalog of fishes . special publication , california academy of sciences , san francisco . 3 vols . 2905 p .

chen , y . y . and x . chu , 1998 . danioninae . p . 19 - 61 . in chen , y . - y . and et al . ( eds ) . fauna sinica . osteichthyes . cypriniformes ii . science press , beijing , 531p .

endo , m . and y . iwatsuki , 1998 . anomalies of wild fishes in the waters of miyazaki , southern japan . bull . faculty agric . 45 ( 1 . 2 ) : 27 - 35 .

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hanel , l . and j . novák , 2002 . ? eské názvy zivo ? ich ? v . ryby a ryboviti obratlovci ( pisces ) 3 . , maloústí ( gonorhynchiformes ) - máloostní ( cypriniformes ) . národní muzeum ( zoologické odd ? lení ) , praha .

jang , m . - h . , j . - g . kim , s . - b . park , k . - s . jeong , g . - i . cho and g . - j . joo , 2002 . the current status of the distribution of introduced fish in large river systems of south korea . internat . rev . hydrobiol . 87 ( 2 - 3 ) : 319 - 328 .

anonymous , 2003 . fish collection of the royal ontario museum . royal ontario museum .

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varjo , m . , l . koli and h . dahlström , 2004 . kalannimiluettelo ( versio 10 / 03 ) . suomen biologian seura vanamo ry .

sado , t . and s . kimura , 2005 . developmental morphology of the indian cyprinid fish barilius canarensis . ichthyol . res . 52 : 360 - 363 .

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the pyrgotidae are an unusual family of flies ( diptera ) , one of only two families of cyclorrhapha that lack ocelli .

most species are " picture-winged " ( i.e , have patterns of bands or spots on the wings ) , as is typical among the tephritoidea , but unlike other tephritoids , they are endoparasitoids ; the females pursue scarab beetles in flight , laying an egg on the beetle 's back under the elytra where the beetle can not reach it .

the egg hatches and the fly larva enters the body cavity of the beetle , feeding and eventually killing the host before pupating .

in the united states , some species of pyrgota and sphecomyiella can be quite common in areas where their host beetles ( typically the genus phyllophaga , or " june beetles " ) are abundant .

like their host beetles , these flies are primarily nocturnal , and are often attracted to artificial lights .

pyrgotidae

pyrgotidae <EOT> in family pyrgotidae that we found in the brisbane area , queensland , australia .

chen , s . h . 1947 . chinese and japanese pyrgotidae . sinensia , 17 , 47 - 74 .

in family pyrgotidae are nocturnal . this may explain why we seldom saw them except in the mating season when they are courting on the tree trunk .

4 . a review of australian pyrgotidae ( diptera ) - by sj paramonov , 1958 , australian journal of zoology 6 ( 1 ) 89 - 138 .

korneyev , v . a . 2012 . revision of the genus pyrgotomyia hendel ( diptera : pyrgotidae ) . african invertebrates , 53 ( 1 ) , 187 - 203 .

hennig , w . 1960 . pyrgotidae de madagascar ( diptera ) . mémoires de l’institut scientifique de madagascar , ser . e , ( 1959 ) , 11 , 321 - 353 .

korneyev , v . a . 2004 . genera of palaearctic pyrgotidae ( diptera , acalyptrata ) , with nomenclatural notes and a key . vestnik zoologii , 38 ( 1 ) , 19 - 46 .

. we determined it is in the family pyrgotidae by its wing veins patterns ( sc vein reaching the wing margin at a sharp right angle ) and its absent of incurved lower fronto - orbital bristles .

hering , e . m . 1940 . acalyptraten aus manchukuo . ( diptera : pyrgotidae , drosophilidae , otitidae ) . arbeiten über morphologische und taxonomische entomologie aus berlin - dahlem , 7 ( 4 ) , 288 - 295 .

hendel , f . 1933 . 36 . pyrgotidae . in : lindner , e . , ed . die fliegen der palaearktischen region . e . schweitzerbart . , stuttgart , 5 ( lfg . 73 ) , 1 - 15 .

korneyev , v . a . 2006 b . a revision of afrotropical species of the eupyrgota ( diptera , pyrgotidae ) : the varipennis and melancholica subgroups of species . vestnik zoologii , 40 ( 2 ) , 115 - 130 .

korneyev , v . a . 2006 a . a revision of afrotropical species of the eupyrgota ( diptera , pyrgotidae ) : the spinifemur group and latipennis subgroup of species . vestnik zoologii , 40 ( 1 ) , 3 - 25 .

korneyev , v . a . 2015 . pyrgotid fl ies assigned to apyrgota . iii . species of afropyrgota gen . n . and tylotrypes ( diptera , pyrgotidae ) . vestnik zoologii , 49 ( 1 ) , 25 - 40 .

soós , a . 1984 . family pyrgotidae . in : soos , a . papp , l . , eds . catalogue of palaearctic diptera . vol . 9 . micropezidae - agromyzidae . akademiai kiado , budapest , 36 - 38 .

* korneyev v . a . monophyly , groundplan and sister - groups in the families pyrgotidae , platystomatidae and tephritidae / / in : third international congress of dipterology . abstract volume . university of guelf , 1994 . p . 112 - 113 .

shi , y . - s . 1996 . pyrgotidae . in : w . xue , c . chao , eds . flies of china , vol . 1 . liaoning science and technology press , shenyang , 575 - 595 [ in chinese ] .

steyskal , g . c . 1980 . 42 . family pyrgotidae . in : crosskey , r . w . , ed . a catalogue of the diptera of the afrotropical region . british museum ( natural history ) , london , 556 - 562 .

kim , s . - k . , han , h . - y . 2001 . a systematic study of the genera adapsilia and parageloemyia in korea ( diptera , tephritoidea , pyrgotidae ) . insecta koreana , 18 ( 3 ) , 255 - 291 .

nartshuk , e . p . , korneyev , v . a . 2005 . data on the fauna of pyrgotidae ( diptera , cyclorrhapha ) of the russian far east . far east entomologist , ( 147 ) , 1 - 10 [ in russian ] .

korneyev v . a . & nartshuk e . p . 77 . fam . pyrgotidae / / keys to insects of far east russia . vol . vi . diptera and fleas . part 3 . vladivostok : dal ' nauka . 2002 : 8 p . ( in russian ) .

korneyev , v . a . 2014 a . pyrgotid fl ies assigned to apyrgota . i . new species and synonyms in eupyrgota ( s . str . ) ( diptera , pyrgotidae ) , with description of a new subgenus . vestnik zoologii , 48 ( 2 ) , 111 - 128 .

korneyev , v . a . 2014 b . pyrgotid fl ies assigned to apyrgota . ii . new species and synonyms in eupyrgota ( s . str . ) ( diptera , pyrgotidae ) , with description of a new subgenus . vestnik zoologii , 48 ( 3 ) , 211 - 220 .

steyskal , g . c . 1977 . family pyrgotidae . in : delfinado , d . , hardy , d . e . , eds . a catalog of the diptera of the oriental region , vol . 3 , suborder cyclorrhapha , ( excluding division aschiza ) . university of hawaii press , honolulu , 37 - 43 .

korneyev , v . a . , nartshuk , e . p . 2004 . 80 . fam . pyrgotidae . in : lehr , p . a . , ed . key to the insects of russian far east . vol . vi . diptera and siphonaptera . pt . 3 . dal’nauka vladivostok , 399 - 408 [ in russian ] .

review of the genus geloemyia ( diptera , pyrgotidae ) , with discussion of its taxonomic position . korneyev , v . a . - species assigned to geloemyia hendel , 1908 , trichempodia malloch , 1930 syn . n . , parageloemyia hendel , 1933 syn . n . , and dicranostira enderlein , 1942 syn . n . are shown to be congeneric . geloemyia is refined to include eight species : geloemyia cheni kim , han & korneyev , sp . n . , geloemyia cockerelli ( malloch , 1930 ) comb . n . ( = trichempodia cockerelli malloch , 1930 ) , geloemyia dorsocentralis ( hering , 1940 ) comb . n . ( = adapsilia dorsocentralis hering , 1940 ) , geloemyia quadriseta hendel , 1933 , geloemyia stylata hendel , 1908 , geloemyia wonjuensis ( kim & han , 2001 ) comb . n . ( = parageloemyia wonjuensis kim & han , 2001 ) from eastern asia , geloemyia namibica sp . n . , from mainland africa ( namibia ) and geloemyia trifasciata ( enderlein , 1942 ) comb . n . ( = trichempodia trifasciata enderlein , 1942 ) from madagascar . geloemyia nigrofasiata hendel , 1933 based on a single male is supposed to be a junior synonym of g . quadriseta hendel , 1933 , based on females , but the synonymy is tentative . a key to species is provided . the genus geloemyia belongs in the tribe pyrgotini forming ( or belonging to ) a basal lineage in the subtribe adapsiliina together with pyrgotomyia hendel , 1914 and porpomastix enderlein , 1942 .

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not much is known about their larvae . it is believed that all the pyrgotid larvae are parasites of adult

we saw those flies near bulimba creek in wishart on early summer oct 2007 . there were a number of them there on a large tree trunk . some of them were courting . the courting pair were facing each other , sometimes with tongue touching .

male and female are a little bit different . they were dark brown in colour with brown patterns on wings .

near those flies we found a batch of insect eggs . could they be the eggs of those flies ? we are not sure . just leave the photo here for further study . ( later we found that those egg - shells look like the

after a few days ( still oct 2007 ) , we also saw those flies in karawatha forest . they seem quite common in this time of the year , but not seen at other time .

csiro , division of entomology , melbourne university press , 2nd edition 1991 , p717 .

2 . on the fly , the interactive atlas and key to australian fly families cd rom - hamilton , j . et al . 2006 . brisbane : cbit & abrs .

3 . northern territory insects , a comprehensive guide cd - graham brown , 2009 .

photographed at turtle river state park , north dakota ( 11 june 2010 ) .

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help again can any1 give me the full title of kulon . allat . kozlem thx

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1 schmalhausen institute of zoology , nas of ukraine , vul . b . khmelnytskogo , 15 , kyiv , 01030 ukraine

enderlein g . 1942 . klassifikation der pyrgotiden . sitzungsberichte der gesellschaft naturforschenden freunde zu berlin , ( 1941 ) 2 , 98 - 134 .

hendel , f . 1908 . acht neue pyrgotinen ( dipt . ) . wiener entomologische zeitung , 27 ( 4 - 5 ) , 145 - 153 .

hendel , f . 1909 . diptera . fam . muscaridae , subfam . pyrgotinae . in : wytsman , p . , ed . genera insectorum , ( 1908 ) , ( fasc . 79 ) , 1 - 33 + 1 taf .

hendel , f . 1934 . ubersicht uber die gattungen der pyrgotiden , nebst beschreibung neuer gattungen und arten . encyclopedie entomologique ( b ) ii . dipt . 7 , 141 - 156 .

malloch , j . r . 1930 . xlii . - exotic muscaridae ( diptera ) . - xxix . the annals and magazine of natural history , 5 ( 10 ) , 465 - 484 .

cite score 2016 : 0 . 35 scimago journal rank ( sjr ) 2016 : 0 . 300 source normalized impact per paper ( snip ) 2016 : 0 . 496

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flies moderately large , with rather slender build and rather long legs . wing 6 - 18 mm long , usually strongly patterned with bars , bands , or reticulation . ( steyskal 1987 )

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to cite this page : myers , p . , r . espinosa , c . s . parr , t . jones , g . s . hammond , and t . a . dewey . 2018 . the animal diversity web ( online ) . accessed at https : / / animaldiversity . org .

disclaimer : the animal diversity web is an educational resource written largely by and for college students . adw doesn ' t cover all species in the world , nor does it include all the latest scientific information about organisms we describe . though we edit our accounts for accuracy , we cannot guarantee all information in those accounts . while adw staff and contributors provide references to books and websites that we believe are reputable , we cannot necessarily endorse the contents of references beyond our control .

this material is based upon work supported by the national science foundation grants drl 0089283 , drl 0628151 , due 0633095 , drl 0918590 , and due 1122742 . additional support has come from the marisla foundation , um college of literature , science , and the arts , museum of zoology , and information and technology services .

1 . verves yu . g . , korneyev v . a . , ponurko ya . b . methodics of collecting of several groups of dipterous insects ( diptera , brachycera ) . actual ' ni problemy biologii [ actual problems of biology ] . kiev : vishcha shkola , 1977 : 15 - 18 . [ in ukrainian ]

2 . korneyev v . a . [ korneev v . a . ] significance of the periphallic complex of flies of family tephritidae for special diagnosis and improving the classification of the family . in : scarlato o . a . , ed . : systematics of diptera ( insecta ) : ecological and morphological principles [ proc . of the 2nd all - union dipterological symposium , woronezh / ramon , sept . 1978 ] . leningrad , zoological institute , 1979 : 38 - 41 . [ in russian , translated in english : new delhi , 1985 : 60 - 64 ]

3 . korneyev v . a . on the fruit fly fauna of the european territory of the ussr . vestnik zoologii , 1982 , no 2 : 83 - 84 . [ in russian ]

4 . korneyev v . a . new fruit fly from the miocene of the northern caucasus . palaeont . zhurnal , 1982 , no 4 : 97 - 98 . [ in russian ]

* korneyev v . a . urophora melanocera hering , a species of tephritid flies ( diptera : tephritidae ) new for the ussr fauna . vestnik zoologii , 1983 , no 2 : 54 . [ in russian ]

* korneyev v . a . xyphosia laticauda mg . , a species of tephritid flies ( diptera : tephritidae ) new for the ussr fauna . vestnik zoologii , 1983 , no 3 : 45 . [ in russian ]

giraudi frauenfeld , a species of tephritid flies ( diptera : ephritidae ) new for the ussr fauna . vestnik zoologii , 1983 , no 3 : 45 . [ in russian ]

* korneyev v . a . chaetorellia loricata ( rondani , 1870 ) = ch . holosericea hendel , 1927 , syn . n . vestnik zoologii , 1983 , no 6 : 45 .

5 . korneyev v . a . tephritid flies ( diptera : tephritidae ) of the middle dniepr territory . kiev , 1983 : 28 pp . a manuscript deposited in ukrainian institute of sci . & techn information [ ukrniinti ] 1 . 12 . 1983 , no 1343 , uk - d83 . [ in russian ]

6 . korneyev v . a . new species of tephritid flies of the genus urophora ( diptera : tephritidae ) from tadjikistan . vestnik zoologii , 1984 , no 1 : 60 - 63 . [ in russian , with english summary ]

7 . korneyev v . a . comparative morphology of aedeagus of tephritidae ( diptera ) . in : scarlato o . a . , ed . : diptera ( insecta ) , their systematics , geographic destribution

and ecology . [ proc . of the 3rd all - union dipterological symposium , kiev / belaya tserkov , sept . 15 - 17 , 1982 ] leningrad : zoological institute , 1984 [ 1983 ] : 67 - 72 . [ in russian , english translation is published , complete data not available ]

8 . verves yu . g . , vlasov i . i . , korneyev v . a . dipterous insects of the families syrphidae , otitidae , platystomatidae , tephritidae and sarcophagidae of zaporozh ' ie region . problems of general and molecular biology . kiev , 1984 . no : 86 - 90 . [ in russian ]

* korneyev v . a . evolution of trophical relationships of tephritidae ( diptera ) . ix congress of all - union entomol . soc . , abstracts . kiev : naukova dumka , 1984 , vol . 1 : 243 . [ in russian ]

9 . volkov b . v . , volkova l . b . , korneyev [ korneev ] v . a . the fruit flies ( diptera : tephritidae ) of voroshilovgrad region . in : scarlato o . a , ed . : diptera ( insecta ) of the fauna of the ussr and their significans in ecosystems [ proc . of the 3rd all - union dipterological symposium , kiev / belaya tserkov , sept . 15 - 17 , 1982 ] . leningrad : zoological institute , 1984 : 27 - 28 . [ in russian ]

* zeigerman a . g . , korneyev v . a . drosophila repleta wollast . , a new species of drosophilidae ( diptera ) in the ussr fauna . vestnik zoologii , 1985 , no 1 : 71 .

* korneyev v . a . tribe terastiomyiini stat . n . ( diptera : tephritidae ) . vestnik zoologii , 1985 , no 2 : 86 .

10 . kameneva e . p . , korneyev v . a . tephritid flies ( diptera : tephritidae ) of the ukrainian ssr fauna . problems of general and molecular biology . kiev , 1985 . no 4 : 67 - 71 . [ in russian ]

11 . verves yu . g . , korneyev v . a . a review of the book : " ' catalogue of palaearctic diptera ' , vol . 9 and 10 . budapest , 1984 " . zoologichesky zhurnal , 1985 , 64 , no 9 : 1436 - 1438 . [ in russian ]

12 . korneyev v . a . tephritid flies ( diptera : tephritidae ) of the middle dnieper territory ( with a review of classifications of the family worldwide ) . abstract of cand . sci . dissertation . kiev state university , 1985 : 19 pp . [ in russian ]

13 . korneyev v . a . new species of the genus urophora ( diptera : tephritidae ) from uzbekistan . vestnik zoologii , 1985 , no 5 : 82 - 84 . [ in russian , english summary ]

14 . korneyev v . a . [ korneev v . a . ] fruit flies of the tribe terelliini hendel , 1927 ( diptera : tephritidae ) of the fauna of the ussr . ent . obozr . , 1985 , 64 , no 3 : 626 - 644 . [ in russian ; english translation in : ent . rev . , wash . 65 , no 1 : 35 - 55 ]

15 . korneyev v . a . contribution to the study of fruit flies ( diptera : tephritidae ) of the yemen arab republic . biol . nauki . 1985 , no 11 : 53 - 56 [ in russian ]

16 . korneyev v . a . , zeigerman a . g . a contribution to the fauna of several families of diptera acalyptrata . i . problems of general and molecular biology . kiev , 1986 . no 5 : 47 - 51 . [ in russian ]

17 . korneyev v . a . on the suprageneric taxonomy of tephritidae . abstr . of the intern . congr . dipterol . , budapest . 1986 : 129 . [ in english ]

18 . korneyev v . a . flies of the families otitidae , platystomatidae and tephritidae ( diptera ) from the zeja state natural reserve . in : nartshuk e . p . , ed . : flies ( diptera ) in ecosystems of south siberia and far east . leningrad : zoological institute , 1986 : 44 - 50 . ( proc . of zool . inst . , leningrad ) [ in russian ]

19 . kameneva e . p . , korneyev v . a . contribution to the fauna of diptera acalyptrata of the middle dnieper region . i . families micropezidae , platystomatidae and otitidae . kiev : editorial board of the journal vestnik zoologii : 7 p . a manuscript deposited in all - union institute of sci . & techn . information [ viniti ] 04 . 01 . 1987 , no 7 - d87 . [ in russian ]

20 . korneyev v . a . , remm e . , elberg k . contribution to the fauna of diptera acalyptrata of the middle dnieper region . ii . families sciomyzidae and lauxaniidae . kiev : editorial board of the journal vestnik zoologii : 13 p . a manuscript deposited in all - union institute of sci . & techn . information [ viniti ] 13 . 03 . 1987 , no 1778 - d87 . [ in russian ]

21 . zeigerman a . g . , korneyev v . a . contribution to the fauna of diptera acalyptrata of the middle dnieper region . iii . families aulacigastridae , asteiidae , camillidae and drosophilidae . kiev : editorial board of the journal vestnik zoologii : 9 p . a manuscript deposited in all - union institute of sci . & techn . information [ viniti ] 05 . 01 . 1987 , no 53 - d87 . [ in russian ]

22 . korneyev v . a . the asparagus fly and its taxonomical position in the family tephritidae ( diptera ) . vestnik zoologii , 1987 no 1 : 39 - 44 . [ in russian ]

23 . korneyev v . a . a revision of the subgenus cerajocera stat . n . of the genus terellia ( diptera , tephritidae ) with description of a new species of fruit fly . zoologichesky zhurnal , 1987 , 67 , no 2 : 237 - 243 . [ in russian ; english summary ]

24 . korneyev v . a . little known species of tephritidae ( diptera ) of the ukrainian fauna . in : savtshenko e . n . , ed . : fauna and biocoenotic associations of insects in ukraine . kiev : naukova dumka , 1987 : 83 - 87 . [ in russian ; english summary ]

25 . basov v . m . , korneyev v . a . exit of tephritid flies ( diptera : tephritidae ) from their larval chambers . problems of general and molecular biology . kiev , 1987 . no 6 : 47 - 51 . [ in russian ]

. in : nartshuk e . p . , ed . : two - wingled

insects : systematics , morphology and ecology . [ proc . of the 4th all - union dipterological symposium , alma - ata , sept . 17 - 19 , 1986 ] . leningrad : zoological institute , 1987 : 49 - 56 . [ in russian ; english translation is published , complete data not available ]

european territory of the ussr . in : nartshuk e . p . : diptera and their importance for husbandry and agriculture . [ proc . of the 4th all - union dipterological symposium , alma - ata , sept . 17 - 19 , 1986 ] . leningrad : zoological institute , 1987 : 36 - 39 . [ in russian ; english translation is published , complete data not available ]

28 . korneyev v . a . tephritid flies of the tribes oedaspidini , aciurini and myopitini ( diptera : tephritidae ) from the soviet far east . new contributions to the systematics of insects of the far east . vladivostok , 1987 : 122 - 129 . [ in russian ]

29 . korneyev v . a . , rogochaya e . g . fruit flies - tephritidae ( trypetidae ) . in : dolin v . g . , ed . : pests of agricultures and forest plantations . 2nd revised edition . vol . 2 . kiev : naukova dumka , 1987 : . [ in russian ]

* kameneva e . p . , korneyev v . a . terellia colon ( meigen , 1826 ) = trypeta varia loew , 1869 , syn . n . vestnik zoologii , 1988 , no 5 : 52 . [ in russian ]

30 . korneyev v . a . new and little known species of tephritid flies of the genus terellia r . - d . from middle asia and transcaucasia . ent . obozr . 1988 , 67 : 871 - 875 . [ in russian ; english summary ; translated in english : ent . rev . , wash . ; complete data not available ]

31 . korneyev v . a . terellia ( cerajocera ) setifera ( diptera : tephritidae ) , a species of tephritid flies new for the ussr . vestnik zoologii , 1989 , no 2 : 75 - 78 . [ in russian ; english summary ]

32 . korneyev v . a . a review of palaearctic species of the genus hendrella ( diptera : tephritidae ) . zoologichesky zhurnal , 1989 , 68 , no 6 : 87 - 92 . [ in russian ; english summary ]

hendel , 1927 = acidiella obscuripennis chen , 1948 , syn . n . ( diptera : tephritidae ) . vestnik zoologii , 1989 , no 4 : 87 .

33 . white i . m . , korneyev v . a . a revision of the western palaearctic species of urophora robineau - desvoidy ( diptera : tephritidae ) . syst . entomol . 1989 , 19 , n 3 : 327 - 374 .

34 . verves yu . g . , zeigerman a . g . , kozitskaya a . m . , korneyev v . a . cyclorrhaphous diptera of kanev vicinities dnieper region . problems of general and molecular biology . kiev , 1989 . no 8 : 17 - 26 . [ in russian ]

35 . korneyev v . a . a new species of the genus chetostoma ( diptera : tephritidae ) from armenia . vestnik zoologii , 1990 , no 1 : 20 - 23 . [ in russian ; english summary ]

* korneyev v . a . terellia sarolensis ( agarval < , > kapoor , 1985 ) comb . n . ( diptera : tephritidae ) . vestnik zoologii , 1990 ( 1 ) : 58 .

36 . korneyev v . a . a review of sphenella and paroxyna series of genera ( diptera : tephritidae : tephritinae ) of the eastern palaearctic . in : insects of mongolia . no 11 . leningrad , 1990 : 395 - 470 . [ in russian ]

37 . korneyev v . a . , kameneva e . p . a new species of the genus tephritis ( diptera : tephritidae ) from kasachstan . zoologichesky zhurnal , 1990 , 69 , no 10 : 138 - 140 . [ in russian ; english summary ]

38 . korneyev v . a . a new species of the genus terellia ( diptera : tephritidae ) from moldavia . vestnik zoologii , 1990 , no 5 : 67 - 69 . [ in russian ; english summary ]

39 . korneyev v . a . new improvements in the suprageneric classification of the family tephritidae . 2nd int . congr . dipterology , august 27 - september 1 , 1990 . bratislava , czechoslovakia . abstract volume : 118 .

40 . korneyev v . a . , kameneva e . p . tephritid flies associated with wild cynarae in the european territory of the u . s . s . r . 2nd int . congr . dipterology , august 27 - september 1 , 1990 . bratislava , czechoslovakia . abstract volume : 119 .

41 . korneyev v . a . tephritid flies of the subfamilies phytalmiinae , acanthonevrinae and adraminae ( diptera : tephritidae ) of the soviet far east . new contributions to the systematics of insects of the far east . vladivostok , 1990 : 116 - 125 . [ in russian ]

42 . korneyev v . a . , white i . m . tephritid flies of the eastern palaearctic species of urophora r . - d . ( diptera : tephritidae ) i . a key to subgenera and a revision of species ( except for the subgenus urophora s . str . ) ent . obozr . 1991 , 70 ( 1 ) : 214 - 228 . [ in russian ; english summary ; english translation expected in entomol . rev . , wash . ]

43 . korneyev v . a . tephritid flies of the genera allied to euleia ( diptera : tephritidae ) in the ussr . communication i . vestnik zoologii , 1991 , no 3 : 8 - 17 . [ in russian ; english summary ]

44 . korneyev v . a . tephritid flies of the genera allied to euleia ( diptera : tephritidae ) in the ussr . communication ii . vestnik zoologii , 1991 , no 4 : 30 - 37 . [ in russian ; english summary ]

45 . korneyev v . a . new species of platystomatidae ( diptera ) from the maritime territory . zoologichesky zhurnal , 1991 , 70 , no 7 : 145 - 148 . [ in russian ; english summary ]

46 . korneyev v . a . , kameneva e . p . tephritid flies ( diptera : tephritidae ) associated with asteraceous plants in eastern europe . problems of general and molecular biology . kiev , 1992 , no 10 : . in ukrainian ; english and russian summary ]

47 . korneyev v . a . reclassification of the palaearctic tephritidae ( diptera ) . communication i . vestnik zoologii , 1992 , no 4 : 31 - 38 . [ in russian ; english summary ]

48 . korneyev v . a . , white i . m . fruit - flies of the eastern palaearctic species of urophora r . - d ( diptera : tephritidae ) . ii . review of species of the subgenus urophora s . str . ( communication 2 ) . ent . obozr . 1992 , 71 ( 3 ) : 688 - 699 . [ in russian ; english summary ; english translation in entomol . rev . , wash . ]

49 . korneyev v . a . , kameneva e . p . on the consortial associations of asteraceae in western tien - shang . ukrain . botan . zhurn . , 1993 , 50 ( 2 ) : 37 - 50 .

50 . korneyev v . a . , white i . m . fruit - flies of the eastern palaearctic species of urophora r . - d ( diptera : tephritidae ) . ii . review of species of the subgenus urophora s . str . ( communication 2 ) . ent . obozr . 1993 , 72 ( 1 ) : 232 - 247 . [ in russian ; english summary ; english translation expected in entomol . rev . , wash . ]

51 . korneyev v . a . new genus and species of tephritid flies ( diptera : tephritidae ) from the russian far east . zoologichesky zhurnal , 1993 , 72 , no 4 : 142 - 144 . [ in russian ; english summary ]

52 . korneyev v . a . a new species of tephritidae ( diptera ) from ukraine . zoologichesky zhurnal , 1993 , 72 , no 4 : 144 - 146 . [ in russian ; english summary ]

53 . korneyev v . a . a revision of palaearctic fruit flies of the genus homoeotricha hering ( diptera tephritidae tephritinae ) / / russian entomol . j . , 1993 . vol . 2 , no . 3 - 4 . p . 119 - 128 .

54 . verves yu . g . , zeigerman a . g . , zrazhevski s . f . , kozitskaya a . m . , korneyev v . a . , tanskaya t . f . class insecta - insects . cyclorrhaphous flies ( diptera , cyclorrhapha ) / / флора и фауна заповедников . вып . 50 . беспозвоночные животные каневского заповедника . [ flora and fauna or natural reserves . issue 50 . invertebrate animals of the kanev natural reserve ] moscow : commission of r [ ussian ] a [ cademy of ] s [ ciences ] for the natural reserve business , institute of evolutionary morphology and ecology of animals of ras , the social and ecological union . 1993 : 12 - 46 . [ in russian ]

55 . korneyev v . a . neoceratitis hendel , a new senior synonym of trirhithromyia hendel ( diptera : tephritidae ) . redescription of the genus and review of species / / j . ukr . ent . soc . , 1993 ( 1994 ) , 1 ( 2 ) : 59 - 62 .

56 . kameneva e . p . , korneyev v . a . holarctic genus pseudoseioptera stackelberg ( diptera : ulidiidae ( = otitidae ) ) . part i . phylogenetic relationships and taxonomic position / / journ . ukr . entomol . soc . , 1993 ( 1994 ) , 1 ( 2 ) : 65 - 72 .

57 . korneyev v . a . reclassification of the palaearctic tephritidae ( diptera ) . communication 2 . vestnik zoologii , 1994 , no 1 : 3 - 17 . [ in russian ; english summary ]

58 . kameneva e . p . , korneyev v . a . holarctic genus pseudoseioptera stackelberg ( diptera : ulidiidae ( = otitidae ) ) . part ii . redescription of the genus and review of species / / j . ukr . ent . soc . , 1993 ( 1995 ) , 1 ( 3 - 4 ) : 69 - 78 .

59 . korneyev v . a . evgeniy nikolaevich savchenko ( 1909 - 1994 ) [ obituary and list of published papers in entomology ] / / j . ukr . ent . soc . , 1993 ( 1995 ) , 1 ( 3 - 4 ) : 3 - 10 . [ in russian ]

60 . korneyev v . a . reclassification of palaearctic tephritidae ( diptera ) . communication 3 . / / vestn . zool . , 1995 , ( 5 - 6 ) : 25 - 48 .

61 . korneyev v . a . new records and synonymy in xyphosiini and tephritini ( diptera : tephritidae : tephritinae ) from the far east russia / / russian entomol . j . , 1995 , 4 ( 1 - 4 ) : 115 - 125 .

62 . korneyev v . a . , merz b . a new species of the genus terellia ( diptera : tephritidae ) from central asia / / j . ukr . ent . soc . , 1994 ( 1995 ) , 2 ( 2 ) : 57 - 60 .

63 . korneyev v . a . identity of vidalia diddiba dirlbek and dirlbekova and v . jibadaua dirlbek and dirlbekova / / int . j . dipterol . res . 1996 , 7 ( 2 ) : 133 - 134 .

64 . korneyev v . a . the status of urophora dzieduszyckii frauenfeld ( diptera : tephritidae : tephritinae : myopitini ) / / ann . naturhist . mus . wien . 1996 , 98b : 525 - 528 .

65 . korneyev v . a . , white i . m . fruit - flies of the eastern palaearctic species of urophora r . - d ( diptera : tephritidae ) . ii . review of species of the subgenus urophora s . str . ( communication 3 ) . / / ent . obozr . 1996 , 75 ( 2 ) : 463 - 477 . [ in russian ; english summary ; english translation expected in entomol . rev . , wash . ]

66 . korneyev v . a . , peck l . v . sem . tephritidae ( trypetidae ) - mukhi - pestrokrylki - fruit - flies . / / genetical fund cadastre of kyrghyzstan . volume iii . superclassis hexapoda ( entognatha and insecta ) . bishkek , 1996 . 296 - 299 .

[ корнеев в . а . , пэк л . в . сем . tephritidae ( trypetidae ) - мухи - пестрокрылки - fruit - flies . / / кадастр генетического фонда кыргызстана . т . iii . надкласс hexapoda - шестиногие ( entognatha и insecta ) . - бишкек , 1996 , с . 296 - 299 . ( а всего том из 406 стр . ) ]

67 . korneyev v . a . , merz b . a new species of rhagoletis loew ( diptera : tephritidae ) , with notes on central asian species / / j . ukr . ent . soc . , 1997 . 3 ( 1 ) : 55 - 64 .

68 . korneyev v . a . a revision of palaearctic campiglossa species assigned to gonioxyna ( diptera : tephritidae ) / / j . ukr . ent . soc . , 1997 . 3 ( 2 ) : 19 - 28 . ( nov 10 1997 )

69 . korneyev v . a . new data and nomenclatural notes on the tephritidae ( diptera ) of far east russia . i / / j . ukr . ent . soc . , 1997 . 3 ( 2 ) : 29 - 35 . ( nov 10 1997 )

* korneyev v . a . the first record of the family odiniidae ( diptera : acalyptrata ) from the ukraine / / vestn . zool . , 1997 , 31 ( 4 ) : 88 .

70 . korneyev v . a . rhagoletis berberidis ( diptera , tephritidae , trypetinae ) , the first faunistic record from ukraine . / / vestn . zool . , 1997 , 31 ( 5 - 6 ) : 94 - 95 [ in ukrainian ; english and russian summary ]

71 . korneyev v . a . new data and nomenclatural notes on the tephritidae ( diptera ) of far east russia . ii / / j . ukr . ent . soc . , 1997 . 3 ( 3 - 4 ) : 5 - 48 .

72 . korneyev v . a . , merz b . a supplement to revision of fruit - flies of the genus urophora r . - d . ( diptera , tephritidae ) of eastern palaearctic . / / ent . obozr . 1998 , 77 ( 2 ) : 512 - 522 . [ in russian ; english summary ; english translation expected in entomol . rev . , wash . ]

* korneyev v . a . the first record of the family carnidae ( diptera : acalyptrata ) from the ukraine / / vestn . zool . , 1999 , 33 ( 1 - 2 ) : 88 .

* korneyev v . a . the first record of the family megamerinidae ( diptera : acalyptrata ) from the ukraine / / vestn . zool . , 1999 , 33 ( 1 - 2 ) : 88 .

73 . korneyev v . a . , verves yu . g . a review of the book " fauna helvetica . 1 . diptera - checklist " / / vestn . zool . , 1999 , 33 ( 1 - 2 ) : 111 .

74 . hernandez ortiz v . , kameneva e . p . & korneyev v . a . a new genus and species of the picture - winged flies ( diptera : ulidiidae : otitinae ) from mexico / / j . ukr . ent . soc . , 1998 [ 1999 ] . 4 ( 1 - 2 ) : 73 - 79 .

75 . korneyev v . a . a review of the book " fauna helvetica . 1 . diptera - checklist " / / j . ukr . ent . soc . , 1998 [ 1999 ] . 4 ( 1 - 2 ) : 80 .

76 . korneyev v . a . , white i . m . fruit - flies of the genus urophora r . - d ( diptera , tephritidae ) of east palaearctic . iii . key to species / / ent . obozr . 1999 , 78 ( 2 ) : 464 - 482 . [ in russian ; english summary ; english translation expected in entomol . rev . , wash . ]

77 . korneyev v . a . phylogenetic relationships among higher groups of the superfamily tephritoidea / / fruit flies ( diptera : tephritidae ) : phylogeny and evolution of behavior . proc . of symposium , xalapa , veracruz , mexico , february 16 - 21 , 1998 . crc press : boca raton , london , new york , washington d . c . 1999 . p . 3 - 22 .

78 . korneyev v . a . phylogenetic relationships among higher groups of tephritidae / / fruit flies ( diptera : tephritidae ) : phylogeny and evolution of behavior . proc . of symposium , xalapa , veracruz , mexico , february 16 - 21 , 1998 . crc press : boca raton , london , new york , washington d . c . 1999 . p . 73 - 114 .

79 . korneyev v . a . phylogeny of tephritinae : relationships of the tribes and subtribes / / fruit flies ( diptera : tephritidae ) : phylogeny and evolution of behavior . proc . of symposium , xalapa , veracruz , mexico , february 16 - 21 , 1998 . crc press : boca raton , london , new york , washington d . c . 1999 . p . 549 - 580 .

80 . korneyev v . a . , white i . m . fruit - flies of the genus urophora r . - d ( diptera , tephritidae ) of east palaearctic . iv . conclusion / / ent . obozr . 2000 , 79 ( 1 ) : 239 - 253 . [ in russian ; english summary ; english translation expected in entomol . rev . , wash . ]

* korneyev v . a . nomenclatural changes in the tephritidae ( diptera ) / / vestn . zool . , 2000 , 34 ( 4 ) : 16 .

81 . korneyev v . a . , dirlbek j . the fruit flies ( diptera : tephritidae ) of syria , jordan and iraq / / studia dipterologica , 2000 , 7 ( 2 ) : 463 - 482 .

82 . korneyev v . a . new records of oriental ctenostylidae ( diptera acalyptrata ) , with discussion of the position of the family / / vestn . zool . , 2001 , 35 ( 3 ) : 47 - 60 .

83 . korneyev v . a . a key to genera of palaearctic platystomatidae ( diptera ) , with descriptions of a new genus and new species / / entomological problems ( bratislava ) , 2001 , 32 ( 1 ) : 1 - 16 .

84 . korneyev v . a . 78 . fam . platystomatidae - signal flies / / keys to insects of far east russia . vol . vi . diptera and fleas . part 2 . vladivostok : dal ' nauka . 2001 : 286 - 295 ( in russian ) .

korneyev v . a . new and little known eurasian dithrycini ( diptera , tephritidae ) / / vestn . zool . , 2002 , 36 ( 3 ) : 13 p .

korneyev v . a . a revision of the quadratula group of the genus terellia r . - d . ( diptera : tephritidae ) / / i internat . tephritidological symposium ; israel , 2000 . proceedings . , 2002 : 16 p .

kameneva e . p . & korneyev v . a . myennidini , a new tribe of the subfamily otitinae ( diptera : ulidiidae ) , with discussion of suprageneric classification of the family / / i internat . tephritidological symposium ; israel , 2000 . proceedings . , 2002 : 96 p .

korneyev v . a . & ovtshinnikova o . g . 79 . fam . tephritidae / / keys to insects of far east russia . vol . vi . diptera and fleas . part 3 . vladivostok : dal ' nauka . 2002 : 92 p . ( in russian ) .

seitz a . , zwoelfer h . & korneyev v . a . tephritidae . / / a . raman , c . w . schaeffer & t . m . withers , eds . biology , ecology and evolution of gall - inducuing arthropods . enfield : science publishers . 2002 : 45 p .

kameneva e . p . & korneyev v . a . family ulidiidae ( = otitidae , pterocallidae ) / / b . brown , f . c . thompson & m . zumbado , ed . manual of central american diptera . 2003 : 4 p .

korneyev v . a . family ctenostylidae ( = lochmostylidae ) / / b . brown , f . c . thompson & m . zumbado , ed . manual of central american diptera . 2003 : 18 p .

korneyev v . a . family platystomatidae ) / / pape , th . , ed . fauna europea : diptera , 2003 : 4 p .

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macroglossum aquila is a moth of the family sphingidae .

it is known from north-eastern india , bangladesh , thailand , southern china , vietnam , malaysia ( peninsular , sarawak ) , indonesia ( sumatra , java , kalimantan ) and the philippines ( luzon ) .

the wingspan is 49 – 54 mm .

macroglossum aquila

macroglossum aquila <EOT> macroglossum aquila boisduval macroglossa aquila boisduval , 1875 , hist . nat . insectes , spec . gen . lep . , 1 : 340 . macroglossum aquila boisduval ; rothschild & jordan , 1903 : 657 .

no one has contributed data records for macroglossum aquila yet . learn how to contribute .

minden pictures stock photos - hummingbird hawk moth ( macroglossum aquila ) hovering near a flower , feeding on the nectar by means of its long probosci . . .

macroglossa aquila boisduval , [ 1875 ] , in : boisduval & guenée , hist . nat . insectes ( spec . gén . lépid . hétérocères ) 1 : 340 . type locality : [ bangladesh , ] silhet [ sylhet ] ; cochinchina [ vietnam ] .

macroglossum geoffmartini ; eitschberger , 2004 , neue ent . nachr . 58 : 10

macroglossum eggeri ; eitschberger , 2004 , neue ent . nachr . 58 : 10

macroglossum haxairei ; eitschberger , 2004 , neue ent . nachr . 58 : 10

macroglossum loeffleri ; eitschberger , 2004 , neue ent . nachr . 58 : 10

macroglossum luteata ; eitschberger , 2004 , neue ent . nachr . 58 : 14

macroglossum oceanicum ; eitschberger , 2004 , neue ent . nachr . 58 : 14

macroglossum perplexum ; eitschberger , 2004 , neue ent . nachr . 58 : 15

macroglossum pseudocorythus ; eitschberger , 2004 , neue ent . nachr . 58 : 15

macroglossum pseudoluteata ; eitschberger , 2004 , neue ent . nachr . 58 : 15

macroglossum stenoxanthum ; eitschberger , 2004 , neue ent . nachr . 58 : 15

macroglossum sulai ; eitschberger , 2004 , neue ent . nachr . 58 : 15

macroglossum corythus fuscicauda ; eitschberger , 2004 , neue ent . nachr . 58 : 12

macroglossum buruensis holland , 1900 ; novit . zool . 7 ( 3 ) : 556 ; tl : buru

macroglossum corythus fulvicaudata ; [ hmw ] ; eitschberger , 2004 , neue ent . nachr . 58 : 12

macroglossum corythus novebudensis ; [ hmw ] ; eitschberger , 2004 , neue ent . nachr . 58 : 12

macroglossum corythus novirlandum ; [ hmw ] ; eitschberger , 2004 , neue ent . nachr . 58 : 12

macroglossum corythus platyxanthum ; [ hmw ] ; eitschberger , 2004 , neue ent . nachr . 58 : 13

macroglossum corythus pylene ; [ hmw ] ; eitschberger , 2004 , neue ent . nachr . 58 : 13

macroglossum corythus xanthurus ; [ hmw ] ; eitschberger , 2004 , neue ent . nachr . 58 : 14

macroglossum sulai eitschberger , 2002 ; neue ent . nachr . 54 : 166 ; tl : sulawesi , selatan , bungku

wingspan : 49 - - 54mm . immediately distinguishable from all other species of macroglossum by the costal edge of hindwing dilated into a rounded lobe in proximal half . forewing upperside with antemedian lines placed more basally than in other species of macroglossum .

macroglossum marquesanum collenette , 1935 ; bull . bishop mus . 114 : 209 ; tl : hivaoa , kaava ridge , 2460ft

? macroglossum palauensis matsumura , 1930 ; trans . sapporo nat . hist . soc . 11 : 120 ; tl : japan

macroglossum eggeri eitschberger , 2002 ; neue ent . nachr . 54 : 173 - 174 ; tl : sulawesi , selatan , parigi

macroglossum pseudocorythus eitschberger , 2002 ; neue ent . nachr . 54 : 158 ; tl : sulawesi , selatan , makki , 800m

macroglossum pseudoluteata eitschberger , 2002 ; neue ent . nachr . 54 : 165 ; tl : sulawesi , selatan , palolo , 800m

macroglossum micacea albibase rothschild , 1905 ; novit . zool . 12 ( 1 ) : 79 ; tl : bougainville i . , solomons

macroglossum perplexum eitschberger , 2002 ; neue ent . nachr . 54 : ? 16 , 170 ; tl : sulawesi , selatan , mamujuju

macroglossum geoffmartini eitschberger , 2002 ; neue ent . nachr . 54 : 172 - 173 ; tl : sulawesi , selatan , n of palopo

macroglossum haxairei eitschberger , 2002 ; neue ent . nachr . 54 : 174 - 175 ; tl : sulawesi , selatan , paredean , 700m

macroglossum sylvia ; hogenes & treadaway , 1998 , nachr . ent . ver . apollo , suppl . 17 : 88 ( part . )

macroglossum fuscicauda rothschild & jordan , 1903 ; novit . zool . 9 ( suppl . ) : 663 ; tl : lifu , loyalty island

macroglossum novebudensis clark , 1926 ; proc . new england zool . cl . 9 : 54 - 55 ; tl : tanna , new hebrides

macroglossum xanthurus rothschild & jordan , 1903 ; novit . zool . 9 ( suppl . ) : 662 ; tl : larat , tenimber island

macroglossum oceanicum rothschild & jordan , 1915 ; novit . zool . 22 ( 2 ) : 292 ; tl : solomon islands , chago archipelago

macroglossum sylvia ; rothschild & jordan , 1903 , 658 ; [ hmw ] ; eitschberger , 2004 , neue ent . nachr . 58 : 15

macroglossum ungues rothschild & jordan , 1903 ; novit . zool . 9 ( suppl . ) : 643 ; tl : buru , amboina , java , etc

macroglossum loeffleri eitschberger , 2002 ; neue ent . nachr . 54 : 171 - 172 ; tl : n . vietnam , 55km nww hanoi , tam dao

macroglossum lepidum rothschild & jordan , 1915 ; novit . zool . 22 ( 2 ) : 287 , pl . 20 , f . 5 ; tl : nias

macroglossum iwasakii matsumura , 1921 ; thous . ins . japan ( additam . ) 4 : 754 , pl . 54 , f . 9 ; tl : japan

macroglossum faro ; rothschild & jordan , 1903 , novit . zool . 9 ( suppl . ) : 665 , pl . 4 , f . 14 ; [ hmw ]

macroglossum sylvia ; inoue , kennett & kitching , 1997 , the moths of thailand 2 : 101 , pl . 30 , f . 136 , p . 146 gen 28

macroglossum kadneri eitschberger , 2004 ; neue ent . nachr . 58 : 7 , 10 , pl . 1 & 2 , f . 5 ; tl : indonesia , sulawesi , selatan , salubai

macroglossum palawana eitschberger & treadaway , 2004 ; neue ent . nachr . 58 : 7 , 10 , pl . 1 & 2 , f . 6 ; tl : philippines , palawan , irawan , 50m

macroglossum trigi eitschberger , 2004 ; neue ent . nachr . 58 : 4 , 11 , pl . 1 & 2 , f . 3 ; tl : indonesia , sulawesi , selatan , n of palopo

macroglossum napolovi eitschberger , 2004 ; neue ent . nachr . 58 : 9 , 10 , pl . 1 & 2 , f . 9 ; tl : n . vietnam , 55km nnw hanoi , tam dao

macroglossum wolframmeyi eitschberger & treadaway , 2004 ; neue ent . nachr . 58 : 8 , 11 , pl . 1 & 2 , f . 8 ; tl : philippines , off palawan , cuyo i . , 400ft

in the male genitalia , uncus gradually narrowed , apically obtuse , slightly downcurved . gnathos rounded apically . valve with stridulatory scales . harpe short , stout , upperside excavated , edges dentate , looking like a brush in lateral view . aedeagus with a patch of teeth near the base of the apical process , which is slender , denticulate basally , along its proximal edge and at the apex , which is obtuse .

china : guangdong ; guangxi ( guilin ; longsheng ) ; hainan ( wanning ) .

northeastern india , bangladesh , thailand , southern china , vietnam , malaysia ( peninsular , sarawak ) , indonesia ( sumatra , java , kalimantan ) , philippines ( luzon ) .

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histoire naturelle des insectes . species général des lépidoptères hétérocères . 1 . sphingides , sésiides , castnides 340

syntypes [ bangladesh : ] silhet [ sylhet ] ; cochinchina [ vietnam ] [ cmnh ] . only a male syntype from " cochinchina " has been found in the cmnh .

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diagnosis . the hindwings have the costa ( seen from below ) with an angular expansion at one third . the yellow band above is constricted centrally by the sharply angular border of the distal black zone . geographical range . n . e . himalaya to sundaland , philippines . habitat preference . two specimens have been taken in recent surveys , both from lowland forest in brunei ( harman , 1981 ) .

macroglossa vialis butler , 1875 ; proc . zool . soc . lond . 1875 : 240 , pl . 36 , f . 5 ; tl : canara

macroglossa interrupta butler , 1875 ; proc . zool . soc . lond . 1875 : 242 , pl . 37 , f . 2 ; tl : darjeeling

macroglossa belia hampson , [ 1893 ] ; fauna br . india ( moths ) 1 : 114 ; tl : ceylon

1024x768 ( ~ 71kb ) oppadake , okinawa , ryukyu , japan , 7 - 93 , photo © s . shuichi haupt

macroglossa calescens butler , 1882 ; ann . mag . nat . hist . ( 5 ) 10 ( 56 ) : 156 ; tl : new britain

macroglossa dohertyi rothschild , 1894 ; novit . zool . 1 ( 1 ) : 67 , pl . 5 , f . 2 ; tl : amboina

sphinx faro cramer , [ 1780 ] ; uitl . kapellen 3 ( 23 - 24 ) : 165

rhamphoschisma godeffroyi butler , 1882 ; ann . mag . nat . hist . ( 5 ) 10 ( 56 ) : 157 ; tl : duke - of - york i .

macroglossa burmanica rothschild , 1894 ; novit . zool . 1 ( 1 ) : 68 , pl . 5 , f . 3 ; tl : burma

heliophila divergens ( walker , 1856 ) ; list spec . lepid . insects colln br . mus . 8 : 94

macroglossa hemichroma butler , 1875 ; proc . zool . soc . lond . 1875 : 243 , pl . 37 , f . 1 ; tl : silhet

indo - australian tropics , australia , hong kong , sumatra , java . see [ maps ]

hong kong , japan , peninsular malaysia , sumatra , borneo , ? sulawesi . see [ maps ]

macroglossa mitchelli ménétriés , 1857 ; cat . lep . petersb . 2 : 95

macroglossa semifasciata hampson , [ 1893 ] ; fauna br . india ( moths ) 1 : 115

macroglossa orientalis butler , 1877 ; trans . zool . soc . lond . 9 ( 19 ) : 528 ; tl : moulmein

naf , seu , ceu , am , ac , s . india , indochina . see [ maps ]

1000x643 ( ~ 116kb ) northern greece , axios delta , june 1995 , photo © dylan lloyd leg .

macroglossa similis rothschild , 1894 ; novit . zool . 1 ( 1 ) : 68 ; tl : oinainissa

macroglossa corythus walker , 1856 ; list spec . lepid . insects colln br . mus . 8 : 92 ; tl : ceylon

macroglossa proxima butler , 1875 ; proc . zool . soc . lond . 1875 : 4 , pl . 1 , f . 1 ; tl : ceylon , canara

macroglossa proxima ; moore , 1882 , lepid . ceylon 2 ( 1 ) : 29 , pl . 91 , f . 1a - b

macroglossa fulvicaudata butler , 1882 ; ann . mag . nat . hist . ( 5 ) 10 ( 56 ) : 155 ; tl : new britain

macroglossa moluccensis rothschild , 1894 ; novit . zool . 1 ( 1 ) : 67 ; tl : batjan , n . molucca islands

macroglossa luteata butler , 1875 ; proc . zool . soc . lond . 1875 : 241 , pl . 37 , f . 5 ; tl : silhet

[ dylan lloyd ] hafanedd , deiniol rd . , bangor , wales , u . k . , ll57 2up

[ maps ] warning ! the maps are automatically generated from the textual information , and the process does not always produce acceptable result ; see about maps for more info .

histoire naturelle des insectes . species général des lépidoptéres hétérocéres . tome premier . sphingides , sésiides , castnides

lepidopterorum amboinensium a dre . l . doleschall annis 1856 - 58 collectorum species novae

reise der österreichischen fregatte novara um die erde in den jahren 1857 , 1858 , 1859 unter den behilfen des commodore b . von wüllerstorf - urbair . zoologischer theil . band 2 . abtheilung 2 . lepidoptera . rhopalocera

sammlung exotischer schmetterlinge , vol . 2 ( [ 1819 ] - [ 1827 ] )

systema naturae per regna tria naturae , secundum clases , ordines , genera , species , cum characteribus , differentiis , symonymis , locis . tomis i . 10th edition

a catalogue of the lepidopterous insects in the museum of the hon . east - india company in horsfield & moore ,

a list of the lepidopterous insects collected by mr . ossian limborg in upper tenasserim , with descriptions of new species

wallengren , 1858 nya fjärilslägten - nova genera lepidopterorum öfvers . vet . akad . förh . 15 : 75 - 84 , 135 - 142 , 209 - 215

if you have corrections , comments or information to add into these pages , just send mail to markku savela keep in mind that the taxonomic information is copied from various sources , and may include many inaccuracies . expert help is welcome .

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this large moth caterpillar gives the threatening appearance of a snake head , which may help to ward off potential predators . the eye - spots are patches of color on the body of the caterpillar , with its real head curled beneath .

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