Action-Oriented Sedan With a Predatory Spirit GIVEN an unforeseen economic slump, an overblown unintended-acceleration drama and an unimaginable earthquake in Japan, it's clear that the recent wounds suffered by Lexus and its parent, Toyota, weren't all self-inflicted. But while external factors and random disasters partly explain how Lexus lost its crown in 2011 - after 11 consecutive years as America's best-selling luxury nameplate - the brand's managers can't merely shake their fists at the heavens. Well before the tsunami, what ailed Lexus (and Toyota) was the calm before the storm: a by-the-book conservatism shaded into complacency, seemingly born of the belief that bulletproof reliability would always carry the day and set these brands apart. That quasi-religious belief, of course, helped Toyota to conquer the world. But as the company now recognizes, such a belief blinded it to customers" desire for cars that speak to the heart as well as the brain. That's especially true for models bought as a splurge. Zestier models from BMW and Mercedes have been dusting Lexus's sales even as Audi, once an underdog, has become the toast of industry tastemakers. Akio Toyoda himself, the race-driving chairman who is a grandson of Toyota's founder, has vowed to restore passion to his brands, which Internet provocateurs have dismissed as "beige." Mr. Toyoda formed a new Global Lexus Division, separate from Toyota, with handpicked leaders who report directly to him. And last August in Pebble Beach, Calif., Mr. Toyoda introduced the redesigned GS 350 sport sedan. He heralded the car's signature "spindle grille" - previewed on the outrageous LF-Gh concept car seen at the New York auto show last year - as "the new face of Lexus." Grimacing and glowering, it's a face that induced déjà vu: where had I seen that mug, and those mandibles, before? Then it hit me: it was in the Central American jungle, in "87. Mr. Toyoda may not realize it, but the new face of Lexus is a dead ringer for the Predator of sci-fi movie fame. Mr. Toyoda has revealed that he initially opposed the styling, but he now thanks his team for fighting him until he gave the project a green light. And there's certainly no denying the passion of a creature, however repulsive it may appear, that skins its victims and keeps their skulls as trophies. Yet the 2013 GS 350 isn't ugly, just mildly odd and alien, and you cannot order one in Slime Green. But because I enjoyed driving the car so much, I declared a truce with the styling. Still, I need to take issue with the huge gaps between the wheel wells and the tires that make the sport sedan appear less hunkered-down than it should. More important, the fourth-generation GS is the most entertaining midrange Lexus sedan in memory. Most unexpectedly, it defies the current midrange trend in which carmakers are prioritizing luxury over performance. Even the BMW 5 Series, the benchmark for action-oriented sedans, has gone softer; the latest model is built on the chassis of the larger 7 Series. How's this for a Hollywood shocker: this Lexus not only has a more overtly aggressive demeanor than the 5 Series, the Audi A6 or the Mercedes E-Class, but its steering also feels more lively and connected. Power is no issue. With 306 horses from a 3.5-liter direct-injection V-6, the rear-drive Lexus manages a 5.7-second sprint from 0 to 60 miles per hour, the company says, in line with the car's main rivals. The all-wheel-drive model is just a bit slower, at 6 seconds. Unlike BMW, Benz and Infiniti, Lexus won't offer a V-8 engine. Instead, the new GS 450h hybrid model will pair a 3.5-liter V-6 with an electric motor and battery for a total of 338 horsepower. A 6-speed transmission does give up one or two gears to its German competitors, though the GS still manages 19 miles per gallon in city driving and 28 on the highway. The all-wheel-drive version is rated 2 m.p.g. less on the freeway; the hybrid leads the lineup with a 29/34 m.p.g. rating. While most of my passengers were ambivalent or tactful about the exterior styling, the contemporary cabin drew universal applause. The interior feels spacious, yet intimate, with expensive-looking surfaces and a tasteful mix of modern and traditional elements. Like its competitors, the Lexus seeks to justify its $50,000-ish price with what I'll call the disconnect of connectivity: the GS wants occupants to feel cocooned in safety and protected from distracted drivers. It has ubiquitous, watchful monitors for blind spots, for drifting out of the lane and for impending collisions. Yet Lexus also insists that technology will free its own drivers to be entertained and informed with no threat to others.