of 22°1, with a maximum of 30. Sixteen flowers were fertilised
with pollen from a long-styled cowslip, and produced 12 cap-
sules, with an average of 24-9 seeds, and a maximum of 42.
Eight flowers were fertilised with pollen from a short-styled
cowslip, but yielded only two capsules, containing 18 and 23
seeds. Hence this plant, in function and partially in structure,
was in an almost exactly intermediate state between the long-
styled and short-styled form, but inclining towards the short-
styled; and this accounts for the low average of seeds which it
produced when spontaneously self-fertilised:

The foregoing five plants thus differ much from one another in
the nature of their fertility. In two individuals a great difference
in the length of the pistil during two succeeding years made no
difference in the number of seeds produced. As all five plants
possessed the male organs of the short-styled form in a perfect
state, and the female organs of the long-styled form in a more
or less complete state, they spontaneously produced a surprising
number of capsules, which generally contained a large average
of remarkably fine seeds. With ordinary cowslips, legitimately
fertilised, I once obtained from plants cultivated in the green-
house the high average, from seven capsules, of 58°7 seeds, with
a@ maximum in one capsule of 87 seeds; but from plants grown
out of doors I never obtained a higher average than 41 seeds.
