KUNIO SEKIYA, IKUO MIURA &amp; MITSUAKI OGATA (2012) A new frog species of the genus &lt;i&gt;Rugosa&lt;/i&gt; from Sado Island, Japan (Anura, Ranidae). &lt;i&gt;Zootaxa&lt;/i&gt;, 3575, 49–62.
Zootaxa