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The last male northern white rhino had been born in Sudan, the last of his kind to be born in the wild.

He was taken to a Czech zoo and then transferred to Kenya in 2009 with the three other remaining fertile northern white rhinos at the time. They were placed under 24-hour armed guard and fed a special diet. "However, despite the fact that they were seen mating, there were no successful pregnancies," the conservancy said.

Rangers caring for Sudan described him as gentle and, as his condition worsened in recent weeks, expressed sadness over his imminent death.

The rhino "significantly contributed to survival of his species as he sired two females," the conservancy said. "Additionally, his geneticsouthern white rhino and another species, the black rhino, are under heavy pressure from poachers who kill them for their horns to supply illegal markets in parts of Asia.

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Roughly 20,000 southern white rhinos remain in Africa. Their numbers dipped below 100 around a century ago, but an intense effort initiated by South African conservationist Ian Player in the mid-20th century turned things around.

This article was reported by The Associated Press.