to the establishment of animal models and animal care directed by QLMY. YQW contributed to the design of the study and the editing of the manuscript. GCW, WLM and QLMY participated in the design of the study and the review of the manuscript. All authors have read and approved the final manuscript.

Acknowledgements
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This work was supported by in part by the National Natural Science Fund of China (30970975, 81171045, 81072875, 31000495 and 31121061) and the Excellent Doctoral Graduate Research Program of Fudan University and the Doctoral Fund of the Ministry of Education of China (20100071110042).
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