As of January 2019, the fleet consists of over 450 W, Z3, A1, B2, C1, C2, D and E class trams, operated from eight depots.

In October 2001, Yarra Trams took delivery of the first of 36 C class Citadis trams which have primarily operated on route 109 to complement the extension to Box Hill.
As part of the acquisition of M>Tram in 2004, Yarra Trams acquired 39 D class Combino trams and oversaw the delivery of another 20 still on order.
They are leased from the Commonwealth Bank, rather than VicTrack.

Heritage W-class trams run solely on the City Circle tourist service.
Due to these trams being limited to 40 km/h (24.9 mph) as a result of a series of modifications due to braking problems, their use is restricted to ensure limited interference with modern trams.

In 2003, Yarra Trams trialled a seating layout which became known as "Apollo".
The trial involved removing 30 seats from a B-class tram and replacing the removed seats with "bum racks" which are found in Citadis trams.
This resulted in a higher capacity for standing passengers on crowded services during peak hours.
A further ten trams received the seating layout during refurbishment.

Ninety E-class are being delivered by Bombardier Transportation, Dandenong.
As of January 2019, the fleet consists of over 450 W, Z3, A1, B2, C1, C2, D and E class trams, operated from eight depots.
In October 2001, Yarra Trams took delivery of the first of 36 C class Citadis trams which have primarily operated on route 109 to complement the extension to Box Hill. As part of the acquisition of M>Tram in 2004, Yarra Trams acquired 39 D class Combino trams and oversaw the delivery of another 20 still on order. They are leased from the Commonwealth Bank, rather than VicTrack.
Heritage W-class trams run solely on the City Circle tourist service. Due to these trams being limited to 40 km/h (24.9 mph) as a result of a series of modifications due to braking problems, their use is restricted to ensure limited interference with modern trams.
In 2003, Yarra Trams trialled a seating layout which became known as "Apollo". The trial involved removing 30 seats from a B-class tram and replacing the removed seats with "bum racks" which are found in Citadis trams. This resulted in a higher capacity for standing passengers on crowded services during peak hours. A further ten trams received the seating layout during refurbishment.
Ninety E-class are being delivered by Bombardier Transportation, Dandenong.
Yarra Trams