The Red Devil (stunt performed by Riley Schmidt and Olga Wilhelmine) is the main antagonist of Season 1.

The Red Devil was the mascot of Wallace University until it was changed due to its association with the killings on campus.
The Devils are both the children of the deceased Sophia Doyle: her son Boone Clemens and her daughter Hester Ulrich (as well as Pete Martinez (who would assist)) who were both raised by Gigi Caldwell/Jess Meyer in a mental asylum after the shocking suicide of their former guardian Amy.
Throughout their time at the institution, the two were taught about murder and orchestrated a 20 year long killing plan against Kappa House.
To not be suspected, the two went under disguises: Hester as a girl with severe scoliosis and had to wear a neck brace whilst Boone pretended to be gay and never enrolled at the school whilst his sister managed to get herself in due to her "neck deformity" and managed to find the costume who they stole of a mascot who was stabbed to death by Gigi.
It is revealed that Hester mostly orchestrated the murders and Boone executed the killings.

On March 25, 2014, Hester and Boone (under the Red Devil costume) broke into Kappa and put hydrochloric acid in Melanie Dorkus' tanning spray which resulted in her being disfigured and the incident being blamed on Chanel Oberlin who rose to power as president and would later be framed by Hester alongside her minions for the Red Devil murders once the plan was complete and throughout the events, carefully put together evidence to frame them successfully.
The killer first makes an appearance when The Red Devil (Boone) appears and kills Chanel #2 before going off to kill Tiffany, Shondell, Coney, Mandy, Dodger, Caulfield, Sam, Jennifer, Earl Grey and a random truck driver, kidnapping Zayday and recruiting Pete (who joined to find out about Hester's identity and killed Roger and a mall cop) to help him kill Gigi who constantly pressured Hester to get rid of her brother due to the sloppy kidnapping incident.
However, Boone was fatally stabbed by Pete (who wanted to finish the murder spree) and died.

After her brother's murder, Hester started to don the costume and decapitated Gigi, stabbed Pete and forced a pizza delivery man to blow up Kappa which resulted in his death.
It is also revealed that Hester turned on the fryer that resulted in Ms Bean's death.
Her identity was revealed to Zayday and Grace who confronted her however she was found with a stiletto in her eye and blamed #5 before later defending herself by hiring a married couple to say that they are her real parents, even her real parents (Sophia Doyle and Wes Gardner) weren't on her record.
The Chanel's were successfully arrested for the murders and sent to the same asylum that Hester and Boone grew up.
Hester, now as the only survivor of the Red Devil team, ends her plan for revenge and the Red Devil conspiracy in January 2016.
At the asylum, Chanel is awoken by a knife wielding Red Devil who was revealed to be her ex-boyfriend Chad Radwell who dresses as the Devil to scare Chanel inside the hospital.
After the Green Meanie killers are killed and everyone's lives return to normal and Chanel is leaving her TV show studio a Red Devil is seen waiting in the backseat of her car.
The Red Devil (stunt performed by Riley Schmidt and Olga Wilhelmine) is the main antagonist of Season 1.
The Red Devil was the mascot of Wallace University until it was changed due to its association with the killings on campus. The Devils are both the children of the deceased Sophia Doyle: her son Boone Clemens and her daughter Hester Ulrich (as well as Pete Martinez (who would assist)) who were both raised by Gigi Caldwell/Jess Meyer in a mental asylum after the shocking suicide of their former guardian Amy. Throughout their time at the institution, the two were taught about murder and orchestrated a 20 year long killing plan against Kappa House. To not be suspected, the two went under disguises: Hester as a girl with severe scoliosis and had to wear a neck brace whilst Boone pretended to be gay and never enrolled at the school whilst his sister managed to get herself in due to her "neck deformity" and managed to find the costume who they stole of a mascot who was stabbed to death by Gigi. It is revealed that Hester mostly orchestrated the murders and Boone executed the killings.
On March 25, 2014, Hester and Boone (under the Red Devil costume) broke into Kappa and put hydrochloric acid in Melanie Dorkus' tanning spray which resulted in her being disfigured and the incident being blamed on Chanel Oberlin who rose to power as president and would later be framed by Hester alongside her minions for the Red Devil murders once the plan was complete and throughout the events, carefully put together evidence to frame them successfully. The killer first makes an appearance when The Red Devil (Boone) appears and kills Chanel #2 before going off to kill Tiffany, Shondell, Coney, Mandy, Dodger, Caulfield, Sam, Jennifer, Earl Grey and a random truck driver, kidnapping Zayday and recruiting Pete (who joined to find out about Hester's identity and killed Roger and a mall cop) to help him kill Gigi who constantly pressured Hester to get rid of her brother due to the sloppy kidnapping incident. However, Boone was fatally stabbed by Pete (who wanted to finish the murder spree) and died.
After her brother's murder, Hester started to don the costume and decapitated Gigi, stabbed Pete and forced a pizza delivery man to blow up Kappa which resulted in his death. It is also revealed that Hester turned on the fryer that resulted in Ms Bean's death. Her identity was revealed to Zayday and Grace who confronted her however she was found with a stiletto in her eye and blamed #5 before later defending herself by hiring a married couple to say that they are her real parents, even her real parents (Sophia Doyle and Wes Gardner) weren't on her record. The Chanel's were successfully arrested for the murders and sent to the same asylum that Hester and Boone grew up. Hester, now as the only survivor of the Red Devil team, ends her plan for revenge and the Red Devil conspiracy in January 2016. At the asylum, Chanel is awoken by a knife wielding Red Devil who was revealed to be her ex-boyfriend Chad Radwell who dresses as the Devil to scare Chanel inside the hospital. After the Green Meanie killers are killed and everyone's lives return to normal and Chanel is leaving her TV show studio a Red Devil is seen waiting in the backseat of her car.
List of Scream Queens characters