Seven units were transferred to London Midland in 2015 to operate the Watford Junction to St Albans service and some peak West Coast Main Line services out of London Euston.
These replaced the seven Class 321 units that transferred to Abellio ScotRail.
In April 2017, one of the five Class 319/4 units, 319455, was swapped for 319433, with the latter receiving a light refresh and a repaint before entering service.

In December 2017, West Midlands Trains (a consortium of Abellio, JR East and Mitsui) took over the operation of the West Midlands rail franchise, with the seven Class 319 units initially leased by London Midland transferring to the new operator under the London Northwestern Railway brand.

A week after the start of the new franchise, London Northwestern Railway leased additional Class 319 units, enabling the company to take their Class 350/1 and 350/3 Desiro units out of service for their planned refurbishment.

London Northwestern Railway's allocation of Class 319s will be replaced during the current franchise by Class 730s.
Seven units were transferred to London Midland in 2015 to operate the Watford Junction to St Albans service and some peak West Coast Main Line services out of London Euston. These replaced the seven Class 321 units that transferred to Abellio ScotRail. In April 2017, one of the five Class 319/4 units, 319455, was swapped for 319433, with the latter receiving a light refresh and a repaint before entering service.
In December 2017, West Midlands Trains (a consortium of Abellio, JR East and Mitsui) took over the operation of the West Midlands rail franchise, with the seven Class 319 units initially leased by London Midland transferring to the new operator under the London Northwestern Railway brand.
A week after the start of the new franchise, London Northwestern Railway leased additional Class 319 units, enabling the company to take their Class 350/1 and 350/3 Desiro units out of service for their planned refurbishment.
London Northwestern Railway's allocation of Class 319s will be replaced during the current franchise by Class 730s.
British Rail Class 319