New Miocene Decapoda (Thalassinidea; Brachyura) from Tierra Del Fuego, Argentina: Paleobiogeographic Implications
ABSTRACT A decapod crustacean fauna of Tierra del Fuego, Argentina, is documented from the middle Miocene Carmen Silva Formation and the early? Miocene Cerro Águila Conglomerate of the Cabo Domingo Group. Three new genera and five new species are named: Asthenognathus australensis, new species; Miotymolus quadratus, new genus and species; Mursia fuegiana, new species; Pharkidodes agele, new genus and species; and Tierrapilumnus edseli, new genus and species. Compilation of all described species of decapods from late Oligocene to early Pliocene exposures in 18 general localities in Patagonia, southern Argentina, and Chile documents two paleobiogeographic provinces, Argentine and Chilean. Coupled with evidence from the biogeographic patterns of associated mollusks, the faunas from Tierra del Fuego have been assigned to the Argentine Paleobiogeographic Province. Comparison of the distribution of Miocene decapods with that of extant decapods (Boschi 2000) leads to the conclusion that the thermal separation of South Atlantic and South Pacific water in the Miocene was more pronounced than today, so that there is no evidence of a discrete Magellanic Biogeographic Province characterizing the high southern latitude region during the Miocene.
MIOCENE FOSSIL DECAPODA (CRUSTACEA: BRACHYURA) FROM PATAGONIA, ARGENTINA, AND THEIR PALEOECOLOGICAL SETTING
ABSTRACT Five previously undescribed decapod taxa have been collected from lower upper Miocene rocks of the Puerto Madryn Formation, Península Valdés region, Chubut Province, Patagonia, Argentina. New species include Osachila valdesensis, Rochinia boschii, Romaleon parspinosus, Panopeus piramidensis, and Ocypode vericoncava. Chaceon peruvianus and Proterocarcinus latus are also reported from the unit, in addition to two indeterminate xanthoid species. Assignment of fossil taxa to genera within the Panopeidae Ortmann, 1893, is difficult due to the marked similarity in dorsal carapace characters among several genera. Panopeus whittenensis Glaessner, 1980, is herein referred to Pakicarcinus Schweitzer et al., 2004. The Puerto Madryn Formation exposed near Puerto Pirámide contains three distinct Facies Associations (1–3), each associated with specific paleoecological and paleoenvironmental conditions, and which recur throughout the section and represent trangressive systems tract (TST) deposits and highstand systems tract (HST) deposits. Within Facies Association 1, near the base of the section at Puerto Pirámide, three paleosurfaces containing invertebrate fossils in life position are exposed and have been carefully mapped in plan view. Because of their sedimentologic, stratigraphic, taphonomic, and paleoecologic features, these three paleosurfaces are considered to be obrution deposits, each preserving a slightly different paleoenvironmental regime in terms of water depth and position with respect to wave base. Paleosurfaces were formed during the waning stages of the transgressive systems tract.
Taphonomy of Isisfordia duncani specimens from the Lower Cretaceous (upper Albian) portion of the Winton Formation, Isisford, central-west Queensland
Taphonomic analysis of fossil material can benefit from including the results of actualistic decay experiments. This is crucial in determining the autochthony or allochthony of fossils of juvenile and adult Isisfordia duncani, a basal eusuchian from the Lower Cretaceous (upper Albian) distal-fluvial-deltaic lower Winton Formation near Isisford. The taphonomic characteristics of the I. duncani fossils were documented using a combination of traditional taphonomic analysis alongside already published actualistic decay data from juvenile Crocodylus porosus carcasses. We found that the I. duncani holotype, paratypes and referred specimens show little signs of weathering and no signs of abrasion. Disarticulated skeletal elements are often found in close proximity to the rest of the otherwise articulated skeleton. The isolated and disarticulated skeletal elements identified, commonly cranial, maxillary and mandibular elements, are typical of lag deposits. The holotype QM F36211 and paratype QM F34642 were classified as autochthonous, and the remaining I. duncani paratypes and referred specimens are parautochthonous. We propose that I. duncani inhabited upper and lower delta plains near the Eromanga Sea in life. Their carcasses were buried in sediment-laden floodwaters in delta plain overbank and distributary channel deposits. Future studies should refer to I. duncani as a brackish water tolerant species.