sentence2_structure: Animal behavior<>can be said<>$$$Animal behavior<>to be controlled<>by genetics and experiences$$$genetics and experiences<>known<>as nature and nurture
sentence1: debate over the extent to which genes (nature) or experiences in a given environment (nurture) control traits such as animal behaviors
sentence2: Animal behavior can be said to be controlled by genetics and experiences, also known as nature and nurture.
question: Animal behavior can be said to be controlled by genetics and experiences, also known as nature and what?