sentence2_structure: The end of a pine tree branch<>bears<>the male cones that produce pollen$$$the male cones<>produce<>pollen
sentence1: The male pine cones, which dangle at the ends of lower branches, have many pollen sacks which produce and hold pollen.
sentence2: The end of a pine tree branch bears the male cones that produce pollen.
question: The end of a pine tree branch bears the male cones that produce what?