sentence2_structure: The cytoskeleton<>holds<>cell organelles<>in place
sentence1: Specific topics include cell metabolism, membrane organization, organelles, compartmentation, membrane trafficking, the cytoskeleton, cell division, and cell signalling.
sentence2: The cytoskeleton holds cell organelles in place within the cytoplasm.
question: What part of the cell holds cell organelles in place within the cytoplasm?