( Vertebrates ( ( have ( ( a ( vertebral column ) ) ( ( in which ) ( ( the notochord ) ( is ( replaced ( by ( ( multiple vertebrae ) ( that ( form ( a backbone ) ) ) ) ) ) ) ) ) ) ) . ) )
(ROOT (S (NP (NNS Vertebrates)) (VP (VBP have) (NP (NP (DT a) (JJ vertebral) (NN column)) (SBAR (WHPP (IN in) (WHNP (WDT which))) (S (NP (DT the) (NN notochord)) (VP (VBZ is) (VP (VBN replaced) (PP (IN by) (NP (NP (JJ multiple) (NN vertebrae)) (SBAR (WHNP (WDT that)) (S (VP (VBP form) (NP (DT a) (NN backbone))))))))))))) (. .)))
Vertebrates have a vertebral column in which the notochord is replaced by multiple vertebrae that form a backbone.
(ROOT (S (NP (NNP Backbone)) (VP (VBZ is) (NP (NP (DT another) (NN name)) (PP (IN for) (NP (DT the) (JJ vertebral) (NN column))))) (. .)))
Backbone is another name for the vertebral column.