( ( debate over ) ( ( the extent ) ( ( to which ) ( ( ( ( genes ( -LRB- ( nature -RRB- ) ) ) or ) ( ( experiences ( in ( a ( given environment ) ) ) ) ( -LRB- ( nurture -RRB- ) ) ) ) ( control ( traits ( such ( as ( animal behaviors ) ) ) ) ) ) ) ) )
(ROOT (S (VP (VB debate) (PRT (RP over)) (NP (NP (DT the) (NN extent)) (SBAR (WHPP (TO to) (WHNP (WDT which))) (S (NP (NP (NP (NNS genes)) (PRN (-LRB- -LRB-) (NP (NN nature)) (-RRB- -RRB-))) (CC or) (NP (NP (NP (NNS experiences)) (PP (IN in) (NP (DT a) (VBN given) (NN environment)))) (PRN (-LRB- -LRB-) (NP (NNP nurture)) (-RRB- -RRB-)))) (VP (VBP control) (NP (NP (NNS traits)) (PP (JJ such) (IN as) (NP (NN animal) (NNS behaviors)))))))))))
debate over the extent to which genes (nature) or experiences in a given environment (nurture) control traits such as animal behaviors
(ROOT (S (NP (NNP Animal) (NN behavior)) (VP (MD can) (VP (VB be) (VP (VBN said) (S (VP (TO to) (VP (VP (VB be) (VP (VBN controlled) (PP (IN by) (NP (NP (NNS genetics) (CC and) (NNS experiences)) (, ,) (VP (ADVP (RB also)) (VBN known) (PP (IN as) (NP (NN nature)))))))) (CC and) (VP (VB nurture)))))))) (. .)))
Animal behavior can be said to be controlled by genetics and experiences, also known as nature and nurture.