( This ( probably ( ( ( ( remains ( true ( despite ( ( ( the fact ) ( that ( nematodes ( as ( ( a group ) ( are ( of ( great ( economic importance ) ) ) ) ) ) ) ) ) ( -LRB- ( ( chiefly ( as ( ( ( parasites and ) pests ) ( of ( both ( ( plants and ) animals ) ) ) ) ) ) -RRB- ) ) ) ) ) ) and ) ( ( are extensively ) ( used ( in ( scientific research ) ) ) ) ) . ) ) )
(ROOT (S (NP (DT This)) (ADVP (RB probably)) (VP (VP (VBZ remains) (ADJP (JJ true) (PP (IN despite) (NP (NP (DT the) (NN fact)) (SBAR (WHNP (WDT that)) (S (VP (VBZ nematodes) (SBAR (IN as) (S (NP (DT a) (NN group)) (VP (VBP are) (PP (IN of) (NP (JJ great) (JJ economic) (NN importance))))))))) (PRN (-LRB- -LRB-) (PP (ADVP (RB chiefly)) (IN as) (NP (NP (NNS parasites) (CC and) (NNS pests)) (PP (IN of) (NP (DT both) (NNS plants) (CC and) (NNS animals))))) (-RRB- -RRB-)))))) (CC and) (VP (VBP are) (ADVP (RB extensively)) (VP (VBN used) (PP (IN in) (NP (JJ scientific) (NN research)))))) (. .)))
This probably remains true despite the fact that nematodes as a group are of great economic importance (chiefly as parasites and pests of both plants and animals) and are extensively used in scientific research.
(ROOT (S (NP (NNS Nematodes)) (VP (MD can) (VP (VB be) (NP (NP (DT a) (NN parasite)) (PP (IN of) (NP (DT both)))))) (. .)))
Nematodes can be a parasite of both.