( ( A prokaryote ) ( ( is ( ( a ( single-celled organism ) ) ( that ( lacks ( ( ( ( ( ( a ( membrane-bound ( nucleus ( -LRB- ( karyon -RRB- ) ) ) ) ) , ) mitochondria ) , ) or ) ( any ( other ( membrane-bound organelles ) ) ) ) ) ) ) ) . ) )
(ROOT (S (NP (DT A) (NN prokaryote)) (VP (VBZ is) (NP (NP (DT a) (JJ single-celled) (NN organism)) (SBAR (WHNP (WDT that)) (S (VP (VBZ lacks) (NP (NP (DT a) (JJ membrane-bound) (NN nucleus) (-LRB- -LRB-) (NN karyon) (-RRB- -RRB-)) (, ,) (NP (NN mitochondria)) (, ,) (CC or) (NP (DT any) (JJ other) (JJ membrane-bound) (NNS organelles)))))))) (. .)))
A prokaryote is a single-celled organism that lacks a membrane-bound nucleus (karyon), mitochondria, or any other membrane-bound organelles.
(ROOT (S (NP (JJ Single-celled) (NNS organisms)) (VP (VBP are) (ADJP (JJ prokaryotes))) (. .)))