( He ( ( ( did ( pioneering research ) ) ( ( using ( ( physical manipulations ) ( of cells ) ) ) ( to ( understand ( ( the mechanisms ) ( of ( ( cytokinesis , ) ( ( the process ) ( ( by which ) ( ( ( a ( cell 's ) ) cytoplasm ) ( is ( divided ( in two ) ) ) ) ) ) ) ) ) ) ) ) ) . ) )
(ROOT (S (NP (PRP He)) (VP (VBD did) (NP (JJ pioneering) (NN research)) (S (VP (VBG using) (NP (NP (JJ physical) (NNS manipulations)) (PP (IN of) (NP (NNS cells)))) (S (VP (TO to) (VP (VB understand) (NP (NP (DT the) (NNS mechanisms)) (PP (IN of) (NP (NP (NNS cytokinesis)) (, ,) (NP (NP (DT the) (NN process)) (SBAR (WHPP (IN by) (WHNP (WDT which))) (S (NP (NP (DT a) (NN cell) (POS 's)) (NN cytoplasm)) (VP (VBZ is) (VP (VBN divided) (PP (IN in) (NP (CD two))))))))))))))))) (. .)))
He did pioneering research using physical manipulations of cells to understand the mechanisms of cytokinesis, the process by which a cell's cytoplasm is divided in two.
(ROOT (S (NP (NNS Cytokinesis)) (VP (VBZ divides) (NP (DT the) (NN cytoplasm)) (PP (IN into) (NP (CD two) (JJ distinctive) (NNS cells)))) (. .)))