( Nebulin ( ( is ( ( a ( giant ( protein molecule ) ) ) ( which ( ( -LRB- ( ( like titin ) -RRB- ) ) ( is ( part ( of ( ( the ( muscle ( cell 's ) ) ) cytoskeleton ) ) ) ) ) ) ) ) ; ) )
(ROOT (S (NP (NNP Nebulin)) (VP (VBZ is) (NP (NP (DT a) (JJ giant) (NN protein) (NN molecule)) (SBAR (WHNP (WDT which)) (S (PRN (-LRB- -LRB-) (PP (IN like) (NP (NN titin))) (-RRB- -RRB-)) (VP (VBZ is) (NP (NP (NN part)) (PP (IN of) (NP (NP (DT the) (NN muscle) (NN cell) (POS 's)) (NN cytoskeleton))))))))) (: ;)))
Nebulin is a giant protein molecule which (like titin) is part of the muscle cell's cytoskeleton;
(ROOT (S (NP (DT The) (ADJP (JJS largest) (VBN known)) (NNS proteins)) (VP (VBP are) (ADJP (JJ titins))) (. .)))