( ( ( ( The ( muscle ( protein titin ) ) ) , ) ( which ( is ( one ( of ( ( the ( key players ) ) ( in muscle ) ) ) ) ) ) ) ( elasticity ( ( ( ( was not ) even ) ( completely ( sequenced ( until ( 1995 ( -LRB- ( ( ( Labeit and ) Kolmerer ) -RRB- ) ) ) ) ) ) ) . ) ) )
(ROOT (S (NP (NP (DT The) (NN muscle) (NN protein) (NN titin)) (, ,) (SBAR (WHNP (WDT which)) (S (VP (VBZ is) (NP (NP (CD one)) (PP (IN of) (NP (NP (DT the) (JJ key) (NNS players)) (PP (IN in) (NP (NN muscle)))))))))) (ADVP (RB elasticity)) (VP (VBD was) (RB not) (ADVP (RB even)) (ADJP (RB completely) (JJ sequenced) (PP (IN until) (NP (NP (CD 1995)) (PRN (-LRB- -LRB-) (NP (NNP Labeit) (CC and) (NNP Kolmerer)) (-RRB- -RRB-)))))) (. .)))
The muscle protein titin, which is one of the key players in muscle elasticity was not even completely sequenced until 1995 (Labeit and Kolmerer).
(ROOT (S (NP (DT The) (ADJP (JJS largest) (VBN known)) (NNS proteins)) (VP (VBP are) (VP (VBN called) (NP (NNS titins)))) (. .)))