( ( ( The migrations ) ( of ( these cells ) ) ) ( ( form ( ( ( ( ( cellular molds ) , ) ( the ( spicule traces ) ) ) , ) ( ( in which ) ( ( ( the cuticle ) ( of ( the spicules ) ) ) ( ( ( is secreted ) , ) ( thus ( determining ( spicule morphology ) ) ) ) ) ) ) ) . ) )
(ROOT (S (NP (NP (DT The) (NNS migrations)) (PP (IN of) (NP (DT these) (NNS cells)))) (VP (VBP form) (NP (NP (JJ cellular) (NNS molds)) (, ,) (NP (DT the) (JJ spicule) (NNS traces)) (, ,) (SBAR (WHPP (IN in) (WHNP (WDT which))) (S (NP (NP (DT the) (NN cuticle)) (PP (IN of) (NP (DT the) (NNS spicules)))) (VP (VBZ is) (ADJP (JJ secreted)) (, ,) (S (ADVP (RB thus)) (VP (VBG determining) (NP (JJ spicule) (NN morphology))))))))) (. .)))
The migrations of these cells form cellular molds, the spicule traces, in which the cuticle of the spicules is secreted, thus determining spicule morphology.
(ROOT (S (NP (NP (DT The) (JJ sharp) (NNS projections)) (SBAR (IN that) (S (NP (JJ specialized) (NNS cells)) (VP (VBP grow))))) (VP (VBP are) (VP (VBN called) (NP (NNS spicules)))) (. .)))