( ( In ( each ( pollen grain ) ) ) ( ( the wall ) ( ( ( ( thickens and ) forms ) ( ( ( ( an ( inner layer ) ) ( -LRB- ( ( the intine ) -RRB- ) ) ) and ) ( ( an ( ( often ( highly sculptured ) ) ( outer layer ) ) ) ( -LRB- ( ( the exine ) -RRB- ) ) ) ) ) . ) ) )
(ROOT (S (PP (IN In) (NP (DT each) (NN pollen) (NN grain))) (NP (DT the) (NN wall)) (VP (VBZ thickens) (CC and) (VBZ forms) (NP (NP (NP (DT an) (JJ inner) (NN layer)) (PRN (-LRB- -LRB-) (NP (DT the) (NN intine)) (-RRB- -RRB-))) (CC and) (NP (NP (DT an) (ADJP (RB often) (RB highly) (JJ sculptured)) (JJ outer) (NN layer)) (PRN (-LRB- -LRB-) (NP (DT the) (NN exine)) (-RRB- -RRB-))))) (. .)))
In each pollen grain the wall thickens and forms an inner layer (the intine) and an often highly sculptured outer layer (the exine).
(ROOT (S (NP (DT Each) (NN pollen) (NN grain)) (VP (VBZ has) (NP (NP (CD two) (NNS coverings)) (: :) (NP (NP (NP (DT the) (NN exine)) (PRN (-LRB- -LRB-) (NP (NP (JJR thicker)) (, ,) (NP (JJ outer) (NN layer))) (-RRB- -RRB-))) (CC and) (NP (DT the) (NN intine))))) (. .)))