( ( ( ( The ( large ( central vacuole ) ) ) ( of ( the ( plant cell ) ) ) ) ( develops ( by ( ( the coalescence ) ( of ( smaller vacuoles ) ) ) ) ) ) ( , ( themselves ( ( derived ( from ( the ( ER ( and ( Golgi apparatus ) ) ) ) ) ) . ) ) ) )
(ROOT (S (S (NP (NP (DT The) (JJ large) (JJ central) (NN vacuole)) (PP (IN of) (NP (DT the) (NN plant) (NN cell)))) (VP (VBZ develops) (PP (IN by) (NP (NP (DT the) (NN coalescence)) (PP (IN of) (NP (JJR smaller) (NNS vacuoles))))))) (, ,) (NP (PRP themselves)) (VP (VBD derived) (PP (IN from) (NP (DT the) (NNP ER) (CC and) (NNP Golgi) (NNP apparatus)))) (. .)))
The large central vacuole of the plant cell develops by the coalescence of smaller vacuoles, themselves derived from the ER and Golgi apparatus.
(ROOT (S (NP (DT The) (NN cell) (NN sap)) (VP (VBZ is) (NP (NP (DT the) (NN liquid)) (PP (IN inside) (NP (DT the) (JJ central) (NN vacuole))))) (. .)))