( Second ( , ( ( the cells ) ( ( are ( ( ( bound together ) ( across ( their cytoplasm ) ) ) ( by ( ( keratin filaments ) ( that ( ( form cell ) ( to ( ( cell connections ) ( -LRB- ( desmosomes -RRB- ) ) ) ) ) ) ) ) ) ) . ) ) ) )
(ROOT (S (ADVP (LS Second)) (, ,) (NP (DT the) (NNS cells)) (VP (VBP are) (VP (VBN bound) (PRT (RP together)) (PP (IN across) (NP (PRP$ their) (NN cytoplasm))) (PP (IN by) (NP (NP (NN keratin) (NNS filaments)) (SBAR (WHNP (WDT that)) (S (VP (VBP form) (NP (NN cell)) (PP (TO to) (NP (NP (NN cell) (NNS connections)) (PRN (-LRB- -LRB-) (NP (NNS desmosomes)) (-RRB- -RRB-))))))))))) (. .)))
Second, the cells are bound together across their cytoplasm by keratin filaments that form cell to cell connections (desmosomes).
(ROOT (S (NP (NNP Plasmodesmata)) (VP (VBP connect) (PP (TO to) (NP (NP (NNS cytoplasms)) (PP (IN in) (NP (DT the) (NN plant) (NN cell)))))) (. .)))