( ( ( Basic kinds ) ( of learning ) ) ( , ( ( studied ( with ( ( rats , ) ( that ( ( have applications ) ( to ( human behavior ) ) ) ) ) ) ) . ) ) )
(ROOT (S (NP (NP (NNP Basic) (NNS kinds)) (PP (IN of) (S (VP (VBG learning))))) (, ,) (VP (VBD studied) (PP (IN with) (NP (NP (NNS rats)) (, ,) (SBAR (WHNP (WDT that)) (S (VP (VBP have) (NP (NNS applications)) (PP (TO to) (NP (JJ human) (NN behavior))))))))) (. .)))
Basic kinds of learning, studied with rats, that have applications to human behavior.
(ROOT (S (NP (JJ Learned) (NN behavior)) (VP (VP (VBZ occurs) (ADVP (RB only)) (PP (IN after) (NP (NN experience) (CC or) (NN practice)))) (CC and) (VP (VBZ describes) (NP (RBS most) (JJ human) (NNS behaviors)))) (. .)))