( ( This species ) ( ( ( feeds primarily ) ( ( ( ( on ( aquatic insects ) ) , ) particularly ) ( on ( caddisflies ( , ( mayflies ( , ( chironomids ( , ( and beetles ) ) ) ) ) ) ) ) ) ) . ) )
(ROOT (S (NP (DT This) (NNS species)) (VP (VBZ feeds) (ADVP (RB primarily)) (PP (PP (IN on) (NP (JJ aquatic) (NNS insects))) (, ,) (RB particularly) (PP (IN on) (NP (NNS caddisflies) (, ,) (NNS mayflies) (, ,) (NNS chironomids) (, ,) (CC and) (NNS beetles))))) (. .)))
This species feeds primarily on aquatic insects, particularly on caddisflies, mayflies, chironomids, and beetles.
(ROOT (S (NP (NNP Overharvesting)) (VP (VBZ is) (NP (DT a) (JJ serious) (NN threat)) (ADVP (RB particularly)) (PP (TO to) (NP (JJ aquatic) (NNS species)))) (. .)))