( ( ( The development ) ( of behavior ) ) ( ( ( pertains ( to ( the ways ) ) ) ( ( ( ( ( in which ) ( behavior ( changes ( over ( ( the lifetime ) ( of ( an animal ) ) ) ) ) ) ) , ) and ) ( how ( ( these changes ) ( are ( affected ( by ( ( ( both genes ) and ) experience ) ) ) ) ) ) ) ) . ) )
(ROOT (S (NP (NP (DT The) (NN development)) (PP (IN of) (NP (NN behavior)))) (VP (VBZ pertains) (PP (TO to) (NP (DT the) (NNS ways))) (SBAR (SBAR (WHPP (IN in) (WHNP (WDT which))) (S (NP (NN behavior)) (VP (VBZ changes) (PP (IN over) (NP (NP (DT the) (NN lifetime)) (PP (IN of) (NP (DT an) (NN animal)))))))) (, ,) (CC and) (SBAR (WHADVP (WRB how)) (S (NP (DT these) (NNS changes)) (VP (VBP are) (VP (VBN affected) (PP (IN by) (NP (NP (DT both) (NNS genes)) (CC and) (NP (NN experience)))))))))) (. .)))
The development of behavior pertains to the ways in which behavior changes over the lifetime of an animal, and how these changes are affected by both genes and experience.
(ROOT (S (NP (NNP Animal) (NNS behaviors)) (VP (VBP are) (VP (VBN controlled) (PP (IN by) (NP (NP (DT both) (NNS genes) (CC and) (NNS experiences)) (PP (IN in) (NP (DT a) (VBN given) (NN envionment))))))) (. .)))