( ( In ( ( the ( charophyte ancestor ) ) ( of ( modern plants ) ) ) ) ( , ( ( ( ( ( ( ( gamete production ) , ) fertilization ) , ) and ) ( development ( of ( the embryo ) ) ) ) ( ( ( were ( highly dependent ) ) ( on ( the ( aquatic environment ) ) ) ) . ) ) ) )
(ROOT (S (PP (IN In) (NP (NP (DT the) (JJ charophyte) (NN ancestor)) (PP (IN of) (NP (JJ modern) (NNS plants))))) (, ,) (NP (NP (JJ gamete) (NN production)) (, ,) (NP (NN fertilization)) (, ,) (CC and) (NP (NP (NN development)) (PP (IN of) (NP (DT the) (NN embryo))))) (VP (VBD were) (ADJP (RB highly) (JJ dependent)) (PP (IN on) (NP (DT the) (JJ aquatic) (NN environment)))) (. .)))
In the charophyte ancestor of modern plants, gamete production, fertilization, and development of the embryo were highly dependent on the aquatic environment.
(ROOT (S (NP (DT The) (NN embryo)) (VP (VBZ develops) (PP (IN in) (NP (NP (DT a) (NN plant)) (PP (IN inside) (NP (DT the) (JJ female) (NN plant))))) (PP (IN after) (NP (NN fertilization)))) (. .)))