( ( ( Two groups ) ( of invertebrates ) ) ( ( have ( ( ( notably ( complex brains ) ) : ) ( ( ( ( arthropods ( -LRB- ( ( ( ( ( ( ( ( insects , ) crustaceans ) , ) arachnids ) , ) and ) others ) -RRB- ) ) ) , ) and ) ( cephalopods ( -LRB- ( ( ( ( ( ( octopuses , ) squids ) , ) and ) ( similar molluscs ) ) -RRB- ) ) ) ) ) ) . ) )
(ROOT (S (NP (NP (CD Two) (NNS groups)) (PP (IN of) (NP (NNS invertebrates)))) (VP (VBP have) (NP (NP (RB notably) (JJ complex) (NNS brains)) (: :) (NP (NP (NP (NNS arthropods)) (PRN (-LRB- -LRB-) (NP (NP (NNS insects)) (, ,) (NP (NNS crustaceans)) (, ,) (NP (NNS arachnids)) (, ,) (CC and) (NP (NNS others))) (-RRB- -RRB-))) (, ,) (CC and) (NP (NP (NNS cephalopods)) (PRN (-LRB- -LRB-) (NP (NP (NNS octopuses)) (, ,) (NP (NNS squids)) (, ,) (CC and) (NP (JJ similar) (NNS molluscs))) (-RRB- -RRB-)))))) (. .)))
Two groups of invertebrates have notably complex brains: arthropods (insects, crustaceans, arachnids, and others), and cephalopods (octopuses, squids, and similar molluscs).
(ROOT (S (SBAR (IN While) (FRAG (ADJP (JJ similar) (PP (TO to) (NP (NNS insects)))))) (, ,) (NP (NNS arachnids)) (VP (VBP are) (NP (NP (JJ eight-legged) (NNS invertebrates)) (SBAR (WHNP (WDT that)) (S (VP (VBP lack) (NP (NN antennae) (CC or) (NNS wings))))))) (. .)))