( ( When ( truncation ( is permitted ) ) ) ( , ( ( uppercase letters ) ( ( represent ( ( the ( ( shortest acceptable ) version ) ) ( of ( the operand ) ) ) ) . ) ) ) )
(ROOT (S (SBAR (WHADVP (WRB When)) (S (NP (NN truncation)) (VP (VBZ is) (VP (VBN permitted))))) (, ,) (NP (JJ uppercase) (NNS letters)) (VP (VBP represent) (NP (NP (DT the) (ADJP (JJS shortest) (JJ acceptable)) (NN version)) (PP (IN of) (NP (DT the) (NN operand))))) (. .)))
When truncation is permitted, uppercase letters represent the shortest acceptable version of the operand.
(ROOT (S (SBAR (WHADVP (WRB When)) (S (VP (VBD represented) (PP (IN by) (NP (DT a) (JJ single) (NN letter)))))) (, ,) (NP (JJ dominant) (NNS alleles)) (VP (VBP are) (VP (VBN represented) (PP (IN by) (NP (JJ uppercase) (NNS letters))))) (. .)))