( ( By ( identifying ( which ( proteins ( bind ( to ( ( specific portions ) ( of titin ) ) ) ) ) ) ) ) ( , ( he ( ( hoped ( to ( correlate ( ( ( ( how ( titin ( fits ( into ( ( the ( overall structure ) ) ( of ( the ( muscle sarcomere ) ) ) ) ) ) ) ) , ) ( as ( well as ) ) ) ( how ( ( these interactions ) ( ( ( coordinate and ) maintain ) ( such ( a ( dynamic system ) ) ) ) ) ) ) ) ) ) . ) ) ) )
(ROOT (S (PP (IN By) (S (VP (VBG identifying) (SBAR (WHNP (WDT which)) (S (NP (NNS proteins)) (VP (VBZ bind) (PP (TO to) (NP (NP (JJ specific) (NNS portions)) (PP (IN of) (NP (NN titin))))))))))) (, ,) (NP (PRP he)) (VP (VBD hoped) (S (VP (TO to) (VP (VB correlate) (SBAR (SBAR (WHADVP (WRB how)) (S (NP (NN titin)) (VP (VBZ fits) (PP (IN into) (NP (NP (DT the) (JJ overall) (NN structure)) (PP (IN of) (NP (DT the) (NN muscle) (NN sarcomere)))))))) (, ,) (CONJP (RB as) (RB well) (IN as)) (SBAR (WHADVP (WRB how)) (S (NP (DT these) (NNS interactions)) (VP (VB coordinate) (CC and) (VB maintain) (NP (PDT such) (DT a) (JJ dynamic) (NN system)))))))))) (. .)))
By identifying which proteins bind to specific portions of titin, he hoped to correlate how titin fits into the overall structure of the muscle sarcomere, as well as how these interactions coordinate and maintain such a dynamic system.
(ROOT (S (NP (DT The) (ADJP (JJS largest) (VBN known)) (NNS proteins)) (VP (VBP are) (ADJP (JJ titins))) (. .)))