( ( ( ( ( ( All ants ) ( rely ( on ( olfaction ( for ( ( their communication ) ( -LRB- ( pheromones -RRB- ) ) ) ) ) ) ) ) , ) and ) ( ( many species ) ( mainly ( use ( ( odor cues ) ( for orientation ) ) ) ) ) ) . )
(ROOT (S (S (NP (DT All) (NNS ants)) (VP (VBP rely) (PP (IN on) (NP (NP (NN olfaction)) (PP (IN for) (NP (NP (PRP$ their) (NN communication)) (PRN (-LRB- -LRB-) (NP (NNS pheromones)) (-RRB- -RRB-)))))))) (, ,) (CC and) (S (NP (JJ many) (NNS species)) (ADVP (RB mainly)) (VP (VBP use) (NP (NP (NN odor) (NNS cues)) (PP (IN for) (NP (NN orientation)))))) (. .)))
All ants rely on olfaction for their communication (pheromones), and many species mainly use odor cues for orientation.
(ROOT (S (NP (NNS Pheromones)) (VP (VBP enable) (NP (NP (NN communication)) (, ,) (SBAR (WHNP (WDT which)) (S (VP (VBZ is) (ADJP (JJ vital) (PP (IN for) (NP (NP (NNS organisms)) (PP (IN like) (NP (NP (NNS ants)) (SBAR (WHNP (WDT that)) (S (VP (VBP live) (PP (IN in) (NP (DT a) (JJ large) (NN community)))))))))))))))) (. .)))