( Telethonin ( ( is ( ( a substrate ) ( of ( ( titin , ) ( ( a ( molecular ruler ) ) ( for ( ( the assembly ) ( of ( ( the sarcomere ) ( by ( providing ( spatially ( defined ( ( binding sites ) ( for ( ( other ( sarcomeric proteins ) ) ( -LRB- ( Gregorio -RRB- ) ) ) ) ) ) ) ) ) ) ) ) ) ) ) ) ) ) . ) )
(ROOT (S (NP (NNP Telethonin)) (VP (VBZ is) (NP (NP (DT a) (NN substrate)) (PP (IN of) (NP (NP (NN titin)) (, ,) (NP (NP (DT a) (JJ molecular) (NN ruler)) (PP (IN for) (NP (NP (DT the) (NN assembly)) (PP (IN of) (NP (NP (DT the) (NN sarcomere)) (PP (IN by) (S (VP (VBG providing) (NP (NP (NN spatially)) (VP (VBN defined) (NP (NP (VBG binding) (NNS sites)) (PP (IN for) (NP (NP (JJ other) (JJ sarcomeric) (NNS proteins)) (PRN (-LRB- -LRB-) (NP (NNP Gregorio)) (-RRB- -RRB-))))))))))))))))))) (. .)))
Telethonin is a substrate of titin, a molecular ruler for the assembly of the sarcomere by providing spatially defined binding sites for other sarcomeric proteins (Gregorio).
(ROOT (S (NP (DT The) (ADJP (JJS largest) (VBN known)) (NNS proteins)) (VP (VBP are) (VP (VBN called) (NP (NNS titins)))) (. .)))