( ( Changes ( in ( ( vaginal ( cytology ( , ( ( behavior and ) endocrinology ) ) ) ) ( throughout ( the ( canine ( estrous ( cycle ( and ( ovulation timing ) ) ) ) ) ) ) ) ) ) ( ( will ( be discussed ) ) . ) )
(ROOT (S (NP (NP (NNS Changes)) (PP (IN in) (NP (NP (JJ vaginal) (NN cytology) (, ,) (NN behavior) (CC and) (NN endocrinology)) (PP (IN throughout) (NP (DT the) (JJ canine) (NNS estrous) (NN cycle) (CC and) (NN ovulation) (NN timing)))))) (VP (MD will) (VP (VB be) (VP (VBN discussed)))) (. .)))
Changes in vaginal cytology, behavior and endocrinology throughout the canine estrous cycle and ovulation timing will be discussed.
(ROOT (S (NP (NNP Ovulation)) (VP (VBZ occurs) (SBAR (IN before) (S (NP (DT the) (NN endometrium)) (VP (VBZ thickens) (PP (IN in) (NP (JJ estrous) (NNS cycles))))))) (. .)))