( ( ( ( ( ( When ( replication ( is successful ) ) ) ( , ( ( the ( new ( HIV virions ) ) ) ( ( ( produce ( quite ( a sparkle ) ) ) , ) though ) ) ) ) , ) and ) ( ( this luminescence ) ( can ( be ( measured ( by ( light sensors ) ) ) ) ) ) ) . )
(ROOT (S (S (SBAR (WHADVP (WRB When)) (S (NP (NN replication)) (VP (VBZ is) (ADJP (JJ successful))))) (, ,) (NP (DT the) (JJ new) (NN HIV) (NNS virions)) (VP (VBP produce) (NP (PDT quite) (DT a) (NN sparkle)) (, ,) (ADVP (RB though)))) (, ,) (CC and) (S (NP (DT this) (NN luminescence)) (VP (MD can) (VP (VB be) (VP (VBN measured) (PP (IN by) (NP (JJ light) (NNS sensors))))))) (. .)))
When replication is successful, the new HIV virions produce quite a sparkle, though, and this luminescence can be measured by light sensors.
(ROOT (S (NP (NP (NNP Producing) (NN light)) (PP (IN without) (NP (NN heat)))) (VP (VBZ is) (VP (VBN called) (S (ADJP (JJ luminescence))))) (. .)))