( ( The mutations ) ( ( were ( ( ( ( either ( complete ( ( deletions or ) deletions ) ) ) ( of ( ( ( one or ) more ) nucleotides ) ) ) , ) ( leading ( to ( frame shifts ) ) ) ) ) . ) )
(ROOT (S (NP (DT The) (NNS mutations)) (VP (VBD were) (NP (NP (DT either) (JJ complete) (NNS deletions) (CC or) (NNS deletions)) (PP (IN of) (NP (QP (CD one) (CC or) (JJR more)) (NNS nucleotides))) (, ,) (VP (VBG leading) (S (VP (TO to) (VP (VB frame) (NP (NNS shifts)))))))) (. .)))
The mutations were either complete deletions or deletions of one or more nucleotides, leading to frame shifts.
(ROOT (S (ADVP (RB Deletions)) (VP (VB remove) (NP (NP (QP (CD one) (CC or) (JJR more)) (NNS nucleotides)) (PP (IN from) (NP (DT the) (NN dna)))))))