( ( Multipolar ( spindles ( and ( lagging chromosomes ) ) ) ) ( ( ( ( are ( frequent ( during ( ( metaphase and ) anaphase ) ) ) ) , ) ( ( suggesting defects ) ( in ( ( the ( mitotic ( apparatus ( and ( chromosome attachment ) ) ) ) ) ( to it ) ) ) ) ) . ) )
(ROOT (S (NP (JJ Multipolar) (NNS spindles) (CC and) (NN lagging) (NNS chromosomes)) (VP (VBP are) (ADJP (JJ frequent) (PP (IN during) (NP (NN metaphase) (CC and) (NN anaphase)))) (, ,) (S (VP (VBG suggesting) (NP (NNS defects)) (PP (IN in) (NP (NP (DT the) (JJ mitotic) (NN apparatus) (CC and) (NN chromosome) (NN attachment)) (PP (TO to) (NP (PRP it)))))))) (. .)))
Multipolar spindles and lagging chromosomes are frequent during metaphase and anaphase, suggesting defects in the mitotic apparatus and chromosome attachment to it.
(ROOT (S (NP (JJ Spindle) (NNS fibers)) (VP (VBP pull) (NP (VBN linked) (NNS chromosomes)) (PP (ADVP (RB apart)) (IN in) (NP (JJ anaphase) (NN i.))))))