( ( Shh gene ) ( ( encodes ( ( ( ( ( a ( secreted ( growth factor ) ) ) , ) ( which ( binds ( to ( its ( cell ( surface receptor ) ) ) ) ) ) ) , ) ( encoded ( by ( the ( patched gene ) ) ) ) ) ) . ) )
(ROOT (S (NP (NNP Shh) (NN gene)) (VP (VBZ encodes) (SBAR (S (NP (NP (DT a) (JJ secreted) (NN growth) (NN factor)) (, ,) (SBAR (WHNP (WDT which)) (S (VP (VBZ binds) (PP (TO to) (NP (PRP$ its) (NN cell) (NN surface) (NN receptor)))))) (, ,)) (VP (VBN encoded) (PP (IN by) (NP (DT the) (JJ patched) (NN gene))))))) (. .)))
Shh gene encodes a secreted growth factor, which binds to its cell surface receptor, encoded by the patched gene.
(ROOT (S (NP (NNS Cells)) (VP (VBP secrete) (VP (VB factor) (NP (DT that) (NN bind)) (PP (TO to) (NP (NNS receptors))))) (. .)))