( ( ( ( ( ( ( ( ( ( The systems ) ( we study ) ) ( range ( from ( small ( RNA catalysts ) ) ) ) ) , ) ( such ( as ( ( ( the hammerhead ) and ) ( hepatitis ( delta ( virus ribozymes ) ) ) ) ) ) ) , ) ( to ( large ( RNA-protein complexes ) ) ) ) , ) ( such ( as ( the spliceosome ) ) ) ) . )
(ROOT (NP (NP (NP (DT The) (NNS systems)) (SBAR (S (NP (PRP we)) (VP (VBP study))))) (ADJP (NN range) (PP (IN from) (NP (JJ small) (NNP RNA) (NNS catalysts)))) (, ,) (PP (JJ such) (IN as) (NP (NP (DT the) (NN hammerhead)) (CC and) (NP (NN hepatitis) (NN delta) (NN virus) (NNS ribozymes)))) (, ,) (PP (TO to) (NP (JJ large) (JJ RNA-protein) (NNS complexes))) (, ,) (PP (JJ such) (IN as) (NP (DT the) (NN spliceosome))) (. .)))
The systems we study range from small RNA catalysts, such as the hammerhead and hepatitis delta virus ribozymes, to large RNA-protein complexes, such as the spliceosome.
(ROOT (S (NP (NP (DT The) (NN discovery)) (PP (IN of) (NP (NNS ribozymes)))) (VP (VBD nullified) (NP (DT the) (NN idea)) (SBAR (IN that) (S (NP (DT all) (JJ biological) (NNS catalysts)) (VP (VBP are) (NP (NNS proteins)))))) (. .)))