( These ( ( can ( be ( used ( both ( on ( ( ( metaphase chromosomes ) and ) ( interphase nuclei ) ) ) ) ) ) ) . ) )
(ROOT (S (NP (DT These)) (VP (MD can) (VP (VB be) (VP (VBN used) (PP (DT both) (IN on) (NP (NP (JJ metaphase) (NNS chromosomes)) (CC and) (NP (JJ interphase) (NNS nuclei))))))) (. .)))
These can be used both on metaphase chromosomes and interphase nuclei.
(ROOT (S (NP (DT The) (NNS chromosomes)) (VP (VB duplicate) (PP (IN in) (NP (DT the) (JJ interphase) (NN phase)))) (. .)))