( Trout ( ( ( can not ) ( ( ( use oxygen ) ( in ( the water ) ) ) ( when ( ( ( metabolic waste ) ( ( -LRB- ( ( ( ( ( ammonia , ) urea ) , ) ( carbon dioxide ) ) -RRB- ) ) concentrations ) ) ( are high ) ) ) ) ) . ) )
(ROOT (S (NP (NNP Trout)) (VP (MD can) (RB not) (VP (VB use) (NP (NN oxygen)) (PP (IN in) (NP (DT the) (NN water))) (SBAR (WHADVP (WRB when)) (S (NP (NP (JJ metabolic) (NN waste)) (PRN (-LRB- -LRB-) (NP (NN ammonia)) (, ,) (NP (NN urea)) (, ,) (NP (NN carbon) (NN dioxide)) (-RRB- -RRB-)) (NNS concentrations)) (VP (VBP are) (ADJP (JJ high))))))) (. .)))
Trout cannot use oxygen in the water when metabolic waste (ammonia, urea, carbon dioxide) concentrations are high.
(ROOT (S (NP (NNS Animals)) (VP (MD can) (VP (VP (VB transport) (NP (NP (DT the) (NN waste) (NN material)) (VP (VBN known) (PP (IN as) (NP (NP (NN urea)) (PP (IN in) (NP (DT the) (JJ circulatory) (NN system)))))))) (CC and) (VP (VB store) (NP (PRP it)) (ADVP (RB safely)) (PP (IN at) (NP (JJ high) (NNS concentrations)))))) (. .)))