( ( ( The ( first stage ) ) ( of photosynthesis ) ) ( ( ( takes place ) ( in ( ( ( the ( thylakoid membrane ) ) and ) ( the ( thylakoid space ) ) ) ) ) . ) )
(ROOT (S (NP (NP (DT The) (JJ first) (NN stage)) (PP (IN of) (NP (NN photosynthesis)))) (VP (VBZ takes) (NP (NN place)) (PP (IN in) (NP (NP (DT the) (JJ thylakoid) (NN membrane)) (CC and) (NP (DT the) (JJ thylakoid) (NN space))))) (. .)))
The first stage of photosynthesis takes place in  the thylakoid membrane and the thylakoid space.
(ROOT (S (NP (DT The) (JJ light) (NNS reactions)) (VP (VBP take) (NP (NP (NN place)) (PP (IN in) (NP (DT the) (NN thylakoid))))) (. .)))