( ( The conifers ) ( ( ( ( are ( the ( ( most successful ) gymnosperms ) ) ) , ) ( especially ( on ( ( dry sites ) ( -LRB- ( ( needles ( are ( drought resistant ) ) ) -RRB- ) ) ) ) ) ) . ) )
(ROOT (S (NP (DT The) (NNS conifers)) (VP (VBP are) (NP (DT the) (ADJP (RBS most) (JJ successful)) (NNS gymnosperms)) (, ,) (PP (RB especially) (IN on) (NP (NP (JJ dry) (NNS sites)) (PRN (-LRB- -LRB-) (S (NP (NNS needles)) (VP (VBP are) (ADJP (RB drought) (JJ resistant)))) (-RRB- -RRB-))))) (. .)))
The conifers  are the most successful gymnosperms, especially on dry sites (needles are drought resistant).
(ROOT (S (NP (NNS Conifers)) (VP (VBP are) (NP (NP (DT the) (ADJP (RBS most) (JJ prevalent)) (NN type)) (PP (IN of) (NP (NNS gymnosperms))))) (. .)))