( ( ( ( ( Radiographic studies ) ( of ( reef ( coral exoskeletons ) ) ) ) : ) ( ( ( rates and ) patterns ) ( of ( coral growth ) ) ) ) . )
(ROOT (NP (NP (NP (JJ Radiographic) (NNS studies)) (PP (IN of) (NP (NN reef) (NN coral) (NNS exoskeletons)))) (: :) (NP (NP (NNS rates) (CC and) (NNS patterns)) (PP (IN of) (NP (NN coral) (NN growth)))) (. .)))
Radiographic studies of reef coral exoskeletons: rates and patterns of coral growth.
(ROOT (S (NP (NNS Corals)) (VP (VBP build) (NP (NP (JJ hard) (NNS exoskeletons)) (SBAR (WHNP (WDT that)) (S (VP (VBP grow) (S (VP (TO to) (VP (VB become) (NP (NN coral) (NNS reefs)))))))))) (. .)))