( ( ( ( Eukaryotes , ) ( which ( include ( ( all ( other organisms ) ) ( except viruses ) ) ) ) ) , ) ( ( have ( ( ( a ( membrane-bound nucleus ) ) ( as ( well as ) ) ) ( membrane-enclosed ( cytoplasmic organelles ) ) ) ) . ) )
(ROOT (S (NP (NP (NNS Eukaryotes)) (, ,) (SBAR (WHNP (WDT which)) (S (VP (VBP include) (NP (NP (DT all) (JJ other) (NNS organisms)) (PP (IN except) (NP (NNS viruses))))))) (, ,)) (VP (VBP have) (NP (NP (DT a) (JJ membrane-bound) (NN nucleus)) (CONJP (RB as) (RB well) (IN as)) (NP (JJ membrane-enclosed) (JJ cytoplasmic) (NNS organelles)))) (. .)))
Eukaryotes, which include all other organisms except viruses, have a membrane-bound nucleus as well as membrane-enclosed cytoplasmic organelles.
(ROOT (S (NP (NNS Eukaryotes)) (VP (VBP have) (NP (NP (NNS nuclei)) (CC and) (NP (ADJP (JJ other)) (ADJP (RB membrane) (VBN bound)) (NNS organelles)))) (. .)))