( ( ( ( Ecology and ) physiology ) ( of ( reef-building corals ) ) ) . )
(ROOT (NP (NP (NN Ecology) (CC and) (NN physiology)) (PP (IN of) (NP (JJ reef-building) (NNS corals))) (. .)))
Ecology and physiology of reef-building corals.
(ROOT (S (NP (NNS Corals)) (VP (VBP build) (NP (NP (JJ hard) (NNS exoskeletons)) (SBAR (WHNP (WDT that)) (S (VP (VBP grow) (S (VP (TO to) (VP (VB become) (NP (NN coral) (NNS reefs)))))))))) (. .)))