( ( While ( genes ( are ( ( influencing ( maternal behavior ) ) ( in mice ) ) ) ) ) ( , ( hormones ( also ( ( have ( an effect ) ) . ) ) ) ) )
(ROOT (S (SBAR (IN While) (S (NP (NNS genes)) (VP (VBP are) (VP (VBG influencing) (NP (JJ maternal) (NN behavior)) (PP (IN in) (NP (NNS mice))))))) (, ,) (NP (NNS hormones)) (ADVP (RB also)) (VP (VBP have) (NP (DT an) (NN effect))) (. .)))
While genes are influencing maternal behavior in mice, hormones also have an effect.
(ROOT (S (NP (NP (NNS Behaviors)) (SBAR (WHNP (WDT that)) (S (VP (VP (VBP are) (ADVP (RB closely)) (VP (VBN controlled) (PP (IN by) (NP (NNS genes))))) (CC and) (VP (VBP have) (NP (NP (RB little)) (PP (TO to) (NP (DT no) (JJ environmental) (NN influence))))))))) (VP (VBP are) (VP (VBN called) (NP (JJ innate) (NNS behaviors)))) (. .)))