( ( Each ( of ( ( the pieces ) ( -LRB- ( ( other ( than pawns ) ) -RRB- ) ) ) ) ) ( ( are ( represented ( by ( an ( uppercase letter ) ) ) ) ) . ) )
(ROOT (S (NP (NP (DT Each)) (PP (IN of) (NP (NP (DT the) (NNS pieces)) (PRN (-LRB- -LRB-) (ADJP (JJ other) (PP (IN than) (NP (NNS pawns)))) (-RRB- -RRB-))))) (VP (VBP are) (VP (VBN represented) (PP (IN by) (NP (DT an) (JJ uppercase) (NN letter))))) (. .)))
Each of the pieces (other than pawns) are represented by an uppercase letter.
(ROOT (S (SBAR (WHADVP (WRB When)) (S (VP (VBD represented) (PP (IN by) (NP (DT a) (JJ single) (NN letter)))))) (, ,) (NP (JJ dominant) (NNS alleles)) (VP (VBP are) (VP (VBN represented) (PP (IN by) (NP (JJ uppercase) (NNS letters))))) (. .)))