( ( model ( of ( ( the timing ) ( of evolution ) ) ) ) ( ( in which ) ( ( evolutionary change ) ( occurs ( at ( a ( ( ( slow and ) steady ) pace ) ) ) ) ) ) )
(ROOT (FRAG (NP (NP (NN model)) (PP (IN of) (NP (NP (DT the) (NN timing)) (PP (IN of) (NP (NN evolution)))))) (SBAR (WHPP (IN in) (WHNP (WDT which))) (S (NP (JJ evolutionary) (NN change)) (VP (VBZ occurs) (PP (IN at) (NP (DT a) (ADJP (JJ slow) (CC and) (JJ steady)) (NN pace))))))))
model of the timing of evolution in which evolutionary change occurs at a slow and steady pace
(ROOT (S (NP (NNP Human) (NN evolution)) (VP (VBZ shows) (SBAR (IN that) (S (NP (JJ evolutionary) (NNS changes)) (ADVP (RB typically)) (VP (VBP occur) (PP (IN at) (NP (DT a) (JJ slow) (NN pace))))))) (. .)))