( specifically ( , ( what ( ( can ( be ( ( changed ( in environments ) ) ( to ( produce ( improved ( ( learning and ) behavior ) ) ) ) ) ) ) . ) ) ) )
(ROOT (S (ADVP (RB specifically)) (, ,) (NP (WDT what)) (VP (MD can) (VP (VB be) (VP (VBN changed) (PP (IN in) (NP (NNS environments))) (S (VP (TO to) (VP (VB produce) (NP (JJ improved) (NN learning) (CC and) (NN behavior)))))))) (. .)))
specifically, what can be changed in environments to produce improved learning and behavior.
(ROOT (S (NP (JJ Learned) (NN behavior)) (VP (VBZ is) (ADJP (JJ adaptive)) (SBAR (IN because) (S (NP (PRP they)) (VP (VBP are) (ADJP (JJ flexible)) (, ,) (S (ADJP (JJ capable) (PP (IN of) (S (VP (VBG changing) (SBAR (IN if) (S (NP (DT the) (NN environment)) (VP (VBZ changes))))))))))))) (. .)))