( ( In ( lin-29 mutants ) ) ( , ( ( spicule development ) ( ( ( ( ( goes awry ) ( after ( ( the generation ) ( of ( spicule cells ) ) ) ) ) , ) ( when ( ( spicule morphogenesis ) ( occurs ( in ( wild-type males ) ) ) ) ) ) . ) ) ) )
(ROOT (S (PP (IN In) (NP (JJ lin-29) (NNS mutants))) (, ,) (NP (JJ spicule) (NN development)) (VP (VBZ goes) (ADVP (RB awry)) (PP (IN after) (NP (NP (DT the) (NN generation)) (PP (IN of) (NP (JJ spicule) (NNS cells))))) (, ,) (SBAR (WHADVP (WRB when)) (S (NP (JJ spicule) (NNS morphogenesis)) (VP (VBZ occurs) (PP (IN in) (NP (JJ wild-type) (NNS males))))))) (. .)))
In lin-29 mutants, spicule development goes awry after the generation of spicule cells, when spicule morphogenesis occurs in wild-type males.
(ROOT (S (NP (NP (DT The) (JJ sharp) (NNS projections)) (SBAR (IN that) (S (NP (JJ specialized) (NNS cells)) (VP (VBP grow))))) (VP (VBP are) (VP (VBN called) (NP (NNS spicules)))) (. .)))