( ( The ( water-splitting reaction ) ) ( ( ( ( occurs ( on ( ( the ( lumenal side ) ) ( of ( the ( thylakoid membrane ) ) ) ) ) ) and ) ( is ( driven ( by ( ( the ( light energy ) ) ( captured ( by ( the photosystems ) ) ) ) ) ) ) ) . ) )
(ROOT (S (NP (DT The) (JJ water-splitting) (NN reaction)) (VP (VP (VBZ occurs) (PP (IN on) (NP (NP (DT the) (JJ lumenal) (NN side)) (PP (IN of) (NP (DT the) (JJ thylakoid) (NN membrane)))))) (CC and) (VP (VBZ is) (VP (VBN driven) (PP (IN by) (NP (NP (DT the) (JJ light) (NN energy)) (VP (VBN captured) (PP (IN by) (NP (DT the) (NNS photosystems))))))))) (. .)))
The water-splitting reaction occurs on the lumenal side of the thylakoid membrane and is driven by the light energy captured by the photosystems.
(ROOT (S (NP (DT The) (JJ light) (NNS reactions)) (VP (VBP take) (NP (NP (NN place)) (PP (IN in) (NP (DT the) (NN thylakoid))))) (. .)))