( ( a ( plant ( `` ( wilts '' ) ) ) ) ( ( when ( water ( is ( lost ( from ( the ( central vacuole ) ) ) ) ) ) ) . ) )
(ROOT (S (NP (DT a) (NN plant) (`` ``) (NN wilts) ('' '')) (SBAR (WHADVP (WRB when)) (S (NP (NN water)) (VP (VBZ is) (VP (VBN lost) (PP (IN from) (NP (DT the) (JJ central) (NN vacuole))))))) (. .)))
a plant &quot;wilts&quot; when water is lost from the central vacuole.
(ROOT (NP (DT The) (JJ large) (JJ central) (NN vacuole) (NN store) (NN water) (. .)))