( ( ( The ( ( strength and ) toughness ) ) ( of cartilage ) ) ( ( is ( provided ( by collagen ) ) ) . ) )
(ROOT (S (NP (NP (DT The) (NN strength) (CC and) (NN toughness)) (PP (IN of) (NP (NN cartilage)))) (VP (VBZ is) (VP (VBN provided) (PP (IN by) (NP (NN collagen))))) (. .)))
The strength and toughness of cartilage is provided by collagen.
(ROOT (S (NP (JJ Lamprey) (NN cartilage)) (VP (VBZ contains) (NP (DT no) (NN collagen))) (. .)))