( ( Because ( ( amphibian reproduction ) ( is ( tightly ( associated ( with ( ( the presence ) ( of water ) ) ) ) ) ) ) ) ( , ( ( ( climatic changes ) ( that ( affect ( water availability ) ) ) ) ( ( ( pose ( a ( particularly ( menacing threat ) ) ) ) ( to ( ( both ( ( aquatic and ) terrestrial-breeding ) ) amphibians ) ) ) . ) ) ) )
(ROOT (S (SBAR (IN Because) (S (NP (NN amphibian) (NN reproduction)) (VP (VBZ is) (ADVP (RB tightly) (S (VP (VBN associated) (PP (IN with) (NP (NP (DT the) (NN presence)) (PP (IN of) (NP (NN water))))))))))) (, ,) (NP (NP (JJ climatic) (NNS changes)) (SBAR (WHNP (WDT that)) (S (VP (VBP affect) (NP (NN water) (NN availability)))))) (VP (VBP pose) (NP (DT a) (RB particularly) (JJ menacing) (NN threat)) (PP (TO to) (NP (ADJP (DT both) (JJ aquatic) (CC and) (JJ terrestrial-breeding)) (NNS amphibians)))) (. .)))
Because amphibian reproduction is tightly associated with the presence of water, climatic changes that affect water availability pose a particularly menacing threat to both aquatic and terrestrial-breeding amphibians.
(ROOT (S (NP (JJ Terrestrial) (NNS biomes)) (VP (VBP are) (VP (VBN associated) (PP (IN with) (NP (NN land))) (, ,) (SBAR (IN while) (S (NP (JJ aquatic) (NNS ones)) (VP (VBP are) (VP (VBN associated) (PP (IN with) (NP (NN water))))))))) (. .)))