( ( ( ( Ecological constraints ) ( on ( modes ( of ( ( ( animal communication ) and ) ( quantitative analysis ) ) ) ) ) ) ( of ( social communication ) ) ) . )
(ROOT (NP (NP (NNP Ecological) (NNS constraints)) (PP (IN on) (NP (NP (NNS modes)) (PP (IN of) (NP (NP (NN animal) (NN communication)) (CC and) (NP (JJ quantitative) (NN analysis)))))) (PP (IN of) (NP (JJ social) (NN communication))) (. .)))
Ecological constraints on modes of animal communication and quantitative analysis of social communication.
(ROOT (S (PP (IN For) (NP (JJ social) (NNS animals))) (, ,) (NP (NN communication)) (VP (VBZ essential)) (. .)))