( ( Emphasis ( on ( ( ( ( ( ( ( membranes , ) ( subcellular ( organelle systems ) ) ) , ) cytoskeleton ) , ) and ) ( cell cycle ) ) ) ) . )
(ROOT (NP (NP (NNS Emphasis)) (PP (IN on) (NP (NP (NNS membranes)) (, ,) (NP (NN subcellular) (NN organelle) (NNS systems)) (, ,) (NP (NN cytoskeleton)) (, ,) (CC and) (NP (NN cell) (NN cycle)))) (. .)))
Emphasis on membranes, subcellular organelle systems, cytoskeleton, and cell cycle.
(ROOT (S (NP (DT The) (NN cytoskeleton)) (VP (VBZ holds) (NP (NP (NN cell) (NNS organelles)) (PP (IN in) (NP (NN place)))) (PP (IN within) (NP (DT the) (NN cytoplasm)))) (. .)))
The cytoskeleton holds cell organelles in place within the cytoplasm.