( ( ( Centrosome-independent ( mitotic ( spindle formation ) ) ) ( in vertebrates ) ) . )
(ROOT (NP (NP (JJ Centrosome-independent) (JJ mitotic) (NN spindle) (NN formation)) (PP (IN in) (NP (NNS vertebrates))) (. .)))
Centrosome-independent mitotic spindle formation in  vertebrates.
(ROOT (S (NP (DT A) (JJ mitotic) (NN spindle)) (VP (VBZ forms) (PP (IN from) (NP (DT the) (NNS centrosomes)))) (. .)))
A mitotic spindle forms from the centrosomes.