( ( ( ( ( The ( male ( pine cones ) ) ) , ) ( which ( dangle ( at ( ( the ends ) ( of ( lower branches ) ) ) ) ) ) ) , ) ( ( have ( ( many ( pollen sacks ) ) ( which ( ( ( produce and ) hold ) pollen ) ) ) ) . ) )
(ROOT (S (NP (NP (DT The) (JJ male) (JJ pine) (NNS cones)) (, ,) (SBAR (WHNP (WDT which)) (S (VP (VBP dangle) (PP (IN at) (NP (NP (DT the) (NNS ends)) (PP (IN of) (NP (JJR lower) (NNS branches)))))))) (, ,)) (VP (VBP have) (NP (NP (JJ many) (NN pollen) (NNS sacks)) (SBAR (WHNP (WDT which)) (S (VP (VBP produce) (CC and) (VBP hold) (NP (NN pollen))))))) (. .)))
The male pine cones, which dangle at the ends of lower branches, have many pollen sacks which produce and hold pollen.
(ROOT (S (NP (NP (DT The) (NN end)) (PP (IN of) (NP (DT a) (JJ pine) (NN tree) (NN branch)))) (VP (VBZ bears) (NP (NP (DT the) (JJ male) (NNS cones)) (SBAR (WHNP (WDT that)) (S (VP (VBP produce) (NP (NN pollen))))))) (. .)))
The end of a pine tree branch bears the male cones that produce pollen.