Colin Howell (bi Ojo Kerinla Osu Keta Odun 1959) je Northern Irish ti a gbe si ewon fun esun apaniyan.. 
ALDH1L2 jẹ́  isoform nínú mitochondrial tí ó fẹ́ fara pé , ALDH1L1, tí ó maa ń sọ 10-formyltetrahydrofolate di African àti carbon dioxide.Àwọn ìtọ́kasíiwe àkàsíwájú sí àyọkà kú.. 