Neuquensaurus

Neuquensaurus (meaning "Neuquén lizard") is a genus of saltasaurid sauropod dinosaur that lived in the Late Cretaceous, about 80 million years ago in Argentina and Uruguay in South America. Its fossils were recovered from outcrops of the Anacleto Formation around Cinco Saltos, near the Neuquén river from which its name is derived.
History.
In 1893, Richard Lydekker named "Titanosaurus australis", based on a series of caudal vertebrae and limb elements. The remains had been found by Santiago Roth and F. Romero in the Neuquén Province of Argentina at the Neuquén River, and were by Lydekker assigned to a single individual. Six caudal vertebrae, with the inventory numbers MLP Ly 1-6-V-28-1, were the holotype of the species. They had probably been found in a layer of the Anacleto Formation.
Some elements that had been referred to "Titanosaurus australis" were reassigned to "Laplatasaurus araukanicus" by Friedrich von Huene in 1929. The same year, von Huene named a "Titanosaurus robustus", and claimed it differed from "T. australis" in the limb material. Von Huene described all the slender limb material to "T. australis", but did not identify any differentiating features between the vertebrae. When describing "T. robustus", von Huene did not really compare other genera and species to it. From the syntype material assigned by von Huene to "T. robustus" José Fernando Bonaparte "et al." in 1978 chose four lectotypes, specimens MLP 26-250, MLP 26-252, MLP 26-254, and MLP 26-259, a left femur, both ulnae, and a left radius.
In 1986, Jaime Eduardo Powell, concluding that "Titanosaurus australis" was less similar than "Laplatasaurus araukanicus" to "Titanosaurus indicus", named a separate genus: "Neuquensaurus". However, he did so in an unpublished dissertation which caused "Neuquensaurus australis" and "Neuquensaurus robustus" to remain invalid "nomina ex dissertatione". In 1990, the two species were assigned to "Saltasaurus" by John Stanton McIntosh, as a "Saltasaurus australis" and a "Saltasaurus robustus", claiming that the features found by Bonaparte were not of sufficient taxonomic importance to justify a generic separation.
In 1992 Powell validly named "Neuquensaurus", with as type species "Titanosaurus australis" of which the "combinatio nova" then is "Neuquensaurus australis". He also found "Titanosaurus robustus" to be assignable to the new genus, but considered it non-diagnostical, and so a "nomen dubium".
Description.
This dinosaur is believed to have possessed armor-like osteoderms. A relatively small sauropod, with a femur only long. It is one of the most completely known of Patagonian sauropods. In addition to the original fossils described by Lydekker in 1893, it is represented by fossils collected in the early twentieth century, and more recent material, including a well preserved and partially articulated specimen described in 2005 (with two associated osteoderms).
Cervical vertebrae are known from multiple specimens of "Neuquensaurus", although few are definitively associated with any other material. The centra are elongate, compressed vertically, and strongly , with an ovoid and rounded neural spine. Diagnostic of Saltasaurinae, the are shortened and robust and the extend far posteriorly to compensate. The lateral projections of the neural arch (transverse processes) are far more robust than in "Isisaurus". As in the cervicals, the dorsal vertebrae bear strongly opisthocoelous centra with lateral pneumatic openings. In the dorsals, however, the pleurocoels are posteriorly pinched and internally subdivided, a regular feature of titanosaurs. The pleurocoels of "Neuquensaurus" are proportionally taller than in "Trigonosaurus" or "Lirainosaurus". A ventral keel is lost throughout the dorsal series of probably 10 vertebrae, although the total count is uncertain. The vertebrae also lean less posteriorly towards the pelvis, initially inclined far beyond their individual centrum. However, they lack the of non-titanosaurs that limit vertebral motion. Two accessory are present in the dorsals of "Neuquensaurus" that are absent even in the close relatives "Saltasaurus" and "Rocasaurus". Unique among all sauropods, the sacrum of "Neuquensaurus" is composed of seven vertebrae. The last vertebra is convex anteriorly and posteriorly, and diagnostically the middle vertebrae are strongly compressed in width. Like other derived titanosaurs, all caudal vertebrae of "Neuquensaurus" are strongly and vertically compressed. There is a prominent ventral depression like in "Saltasaurus" and "Rocasaurus", but contrasting the other genera there is no ridge dividing this depression. Additionally, the neural spines of vertebrae become progressively more reclined, and a prespinal lamina is present in all vertebrae.
"Neuquensaurus" has an almost complete skeleton. Among the preserved elements are a scapula. The scapula is co-ossified with the coracoid, a feature also found in "Opisthocoelicaudia". In general aspect, the scapula resembles "Saltasaurus", "Opisthocoelicaudia", "Lirainosaurus", and "Alamosaurus". The coracoids of the scapulae are roughly quadrangular in shape, and are alike "Saltasaurus" and "Lirainosaurus", but not "Opisthocoelicaudia", "Rapetosaurus", or "Isisaurus", all of which have a rounded coracoid.
Many characters distinguish "Neuquensaurus" from other titanosaurians. The features found by Otero in 2010 are: the possession of posterior caudal centra that are dorsoventrally flattened, and strongly developed fibular lateral tuberosity.
Classification.
"Neuquensaurus" is a derived saltasaurine. Its closest relatives were "Saltasaurus", "Rocasaurus", and "Bonatitan", and together they made up Saltasaurinae.
Paleoecology.
"Neuquensaurus" is known from fossils from the Anacleto Formation.