Tapejara wellnhoferi

Tapejara (from a Tupi word meaning "the old being") is a genus of Brazilian pterosaur from the Cretaceous Period (Santana Formation, dating to about 112 million years ago). "Tapejara" crests consisted of a semicircular crest over the snout, and a bony prong which extended back behind the head.
Species and classification.
The type species and only one currently recognized as valid by most researchers, is "T. wellnhoferi". The specific name honors German paleontologist Peter Wellnhofer. Two larger species, originally named "Tapejara imperator" and "Tapejara navigans", were later classified in the genus "Tapejara". However, several studies have shown that "T. imperator" and "T. navigans" are significantly different from "T. wellnhoferi" and therefore were reclassified into new genera. The species "T. imperator" was given its own genus, "Tupandactylus", by Alexander Kellner and Diogenes de Almeida Campos. Unwin and Martill found that "T. imperator" and "T. navigans" belong in the same genus, and named them "Ingridia imperator" and "I. navigans", respectively. This genus name honored Wellnhofer's late wife Ingrid. 
Because "Tupandactylus" was named first, it retained priority over the name "Ingridia". To complicate matters, both the name "Tupandactylus" and "Ingridia" used the former "Tapejara imperator" as their type species. The scientists who described "Tupandactylus" did not name a "Tupandactylus navigans" (but instead suggested it was synonymous to "Tupandactylus imperator"), and "Tapejara navigans" was not formally reclassified as a distinct species of "Tupandactylus" until 2011.
The cladogram below follows a phylogenetic analysis by Kellner, the describer of "Tapejara", and colleagues in 2019. They recovered "Tapejara" within the Tapejarini (a tribe within the family Tapejaridae), sister taxon to three other genera: "Europejara", "Caiuajara", and "Tupandactylus".
Paleobiology.
Comparisons between the scleral rings of "Tapejara" and modern birds and reptiles suggest that it may have been cathemeral, active throughout the day at short intervals.