Cuspilongus

Cuspilongus is an extinct genus of parasitic wasp in the sawfly family Cephidae. At the time of its description the new genus comprised a single species named Cuspilongus cachecreekensis, known from an Eocene fossil found in North America. Subsequently, Kopylov & Rasnitsyn (2016) transferred the Early Cretaceous (Aptian) species "Mesocephus ghilarovi" Rasnitsyn (1988), known from the Bon-Tsagan locality in Mongolia, to the genus "Cuspilongus" as well.
History and classification.
"C. cachecreekensis" is known only from one fossil, a part and counterpart holotype, specimen number F-1545 & F-1546, which is housed in the collections of the Thompson Rivers University in Kamloops, British Columbia. "Cuspilongus" was described from a specimen which was recovered from outcrops of the early Eocene, Ypresian McAbee Fossil Beds near Cache Creek, British Columbia. The unnamed formation outcropping at the McAbee Fossil Beds preserve an upland temperate flora that was first interpreted as being Microthermal, although further study has shown them to be more mesothermal in nature. The plant community preserved in the McAbee Fossil Beds site is mostly broadleaf pollen with alder and elm dominating, and may represent a successional forest involving multiple volcanic ash eruptions.
"C. cachecreekensis" was first studied by the paleoentomologists S. Bruce Archibald from Simon Fraser University in Burnaby, British Columbia and Alexandr Rasnitsyn of the A. A. Borissiak Paleontological Institute. Their 2015 type description of the new genus and species was published in the journal "Canadian Entomologist". The genus name "Cuspilongus" was coined by the researchers as a combination of the Latin words "cuspis" which means "lance" and "longus" meaning long, in reference to the notable length of the holotype's ovipositor. The specific epithet "cachecreekensis" is in honor of the town of Cache Creek, British Columbia, west of the type locality.
"Cuspilongus cachecreekensis" was one of three sawfly species described in Archibald & Rasnitsyn's 2015 paper, the other two being "Ulteramus republicensis" and "Ypresiosirex orthosemos", from the Klondike Mountain Formation and the McAbee fossil beds respectively.
Description.
The single described female of "C. cachecreekensis" is mostly complete, missing the legs entirely, with the antennae preserved in disarticulated segments, and portions of the abdomen are obscured. The female has a body length of approximately and an overall preserved length of with the ovipositor included. The head and thorax are dark in coloration, while the abdomen is light colored in the preserved and visible areas. Similarly the wings are lightly colored to hyaline, with the exception of the intercostal space, which is notably darkened. The abdomen is damaged but the ovipositor is preserved well, being about long with part of the sheath preserved, with both the ovipositor and sheath showing a downward curve. The genus is distinct from other members of the family in the length of the ovipositor, which is similar to the length of the forewing. Living members of the family have ovipositors that are only up to half the length of the forewing.