Dubreuillosaurus

Dubreuillosaurus is a genus of carnivorous dinosaur from the middle Jurassic Period. It is a megalosaurid theropod. Its fossils were found in France. The only named species, Dubreuillosaurus valesdunensis, was originally described as a species of "Poekilopleuron", "Poekilopleuron? valesdunensis", which is still formally the type species of the genus. It was later renamed "Dubreuillosaurus valesdunensis" when, in 2005, Allain came to the conclusion that it was not part of the genus "Poekilopleuron". Its type specimen, MNHN 1998-13, is only rivalled in the number of preserved elements in this group by that of "Eustreptospondylus". "Dubreuillosaurus" is considered to be the sister species of "Magnosaurus". It did not show signs of insular dwarfism even though it was uncovered on an island.
Discovery and naming.
In 1994, the mayor of Conteville in Normandy, André Dubreuil, noted that during land rehabilitation of the old quarry of "Pierre de Caen", dinosaur bones had surfaced. He secured a partial skull and some ribs and notified the Paris "Muséum national d'Histoire naturelle". However, professional excavations only started in 1998 when the rock of the collapsed quarry face had already been spread by a bulldozer over a considerable surface. It proved necessary to dig up and sieve many cubic metres of rubble over several years, ultimately salvaging about two thousand bone fragments, varying in size between one and ten centimetres. From these scraps a more complete skeleton had to be reassembled.
In 2002, when this process had not yet been completed, Ronan Allain, after having dedicated a thesis to it, named the find as a new species of "Poekilopleuron": "Poekilopleuron? valesdunensis". The specific name referred to the nearby ancient battlefield of the Battle of Val-ès-Dunes, where William the Conqueror had in 1046 defeated his enemies. The naming paper also contained a description of the skull. The question mark after the generic name already indicated the assignment to "Poekilopleuron" was tentative. By 2005, Allain had come to the conclusion that the new species was not part of "Poekilopleuron". He therefore created the new genus name "Dubreuillosaurus" for this species, the name honouring the Dubreuil family. The type species of the genus is the original "Poekilopleuron valesdunensis", the "combinatio nova" is "Dubreuillosaurus valesdunensis". The 2005 paper also contained a description of the postcranial skeleton.
The holotype, MNHN 1998-13, has been uncovered in a layer of the Calcaire de Caen dating from the middle Bathonian. It consists of a fragmentary skeleton with skull. Preserved parts include: the majority of the skull; the splenial and angular of the lower jaw, a natural mold of the surangular, two cervical vertebrae, cervical ribs, seven dorsals, ribs, gastralia, three sacrals, ten caudals, a chevron, a piece of the scapula, a claw of the hand, a partial thighbone, the upper part of a shinbone, a partial fibula, a fifth metatarsal and the first phalanx of the third toe. Considering the incompleteness of most other megalosaurids, this genus is exceptional in having a significant percentage of the skeleton been found. The type specimen of "Dubreuillosaurus" is in the number of preserved elements only rivalled in this group by that of "Eustreptospondylus". The holotype represents a subadult individual.
Description.
"Dubreuillosaurus" was physically comparable to "Eustreptospondylus". The holotype was an individual by Gregory S. Paul estimated to measure long and to weigh . However, this was a subadult; the adult length is uncertain, though it has been indicated as high as .
In 2005, Allain established some distinctive traits. "Dubreuillosaurus" has an unusually low and long skull, with a length three times the height. In the upper corner of the front edge of the nasal branch of the maxilla, a kink is present, separating a convex curvature below from a concave curvature above. The descending branch of the postorbital has a U-shaped cross-section. The parietal bones are not visible in side view. The supratemporal fenestra has a straight inner edge. A paraquadratic fenestra is lacking. A well-developed process is present on the underside of the jugal branch of the ectopterygoid. The rear edge of the jugal branch of the ectopterygoid, in front of the infratemporal fenestra, is deeply grooved. The lower jaw has a large external mandibular fenestra. The "foramen mylohyoideum" is largely oriented to the front and below. The head of the thighbone is directed to the inside and below. The front underside of the thighbone has a concave surface. In 2012 Matthew Carrano added one autapomorphy: on the rear of the braincase there is a notch between the basioccipital and the bone surface formed by the exoccipitals and the opisthotic.
"Dubreuillosaurus" seemed to lack any sort of crest or horns, but the only known specimen is a juvenile, and it is possible that these structures developed in later life. Like its relatives, "Dubreuillosaurus" probably had short, powerful arms with three-fingered hands.
Phylogeny.
In 2002, "Poekilopleuron? valesdunensis" was assigned to the Megalosauridae. In 2012, Carrano "et al." refined this to the Afrovenatorinae, in which it would be the sister species of "Magnosaurus" within the clade.
The phylogenetic position of "Dubreuillosaurus" according to Carrano "et al." is shown by this cladogram:
Paleobiology.
During the Middle Jurassic much of Europe consisted of a number of islands. The fossil of "Dubreuillosaurus" was discovered in sedimentary rocks that were laid down in coastal, mangrove swamps at the east coast of the Armorican Massif. This suggests that it might have hunted fish and other marine prey.
Allain in 2002 described "P? valesdunensis" as a fish eater or piscivore but this was influenced by the fact that the holotype of "Poekilopleuron bucklandii" had been found together with fish remains. Though uncovered at an island "Dubreuillosaurus" did not show the effects of insular dwarfism; like with "Eustreptospondylus" its small size is due to its being a subadult.