Donatia

Donatia is a genus of two cushion plant species in the family Stylidiaceae. The name commemorates Vitaliano Donati, an Italian botanist.
"Donatia" has been placed in the subfamily Donatioideae, described by Johannes Mildbraed in his 1908 taxonomic monograph of the family Stylidiaceae. The subfamily was created to distinguish the difference between the single genus "Donatia" from the five typical genera of the Stylidiaceae that Mildbraed placed in the Stylidioideae subfamily. The subfamily taxonomy represented the taxonomic uncertainty of "Donatia", which had at one point also been placed in the Saxifragaceae. "Donatia" differs sufficiently from the genera in the Stylidiaceae in that it has free stamens and petals, paracytic stomata, and a pollen morphology distinct from the other genera. Because of this and the recent phylogenetic analysis based on "rbcL" genes, more recent treatments have segregated "Donatia" into its own family, the Donatiaceae. The molecular phylogenetic analysis has placed "Donatia" as a sister-group to Stylidiaceae, thus leaving the Stylidiaceae as a monophyletic family.
As early as three years after Mildbraed's publication of the subfamily Donatioideae, other authors began to question the placement and argued for recognition of Donatiaceae. In 1915, Carl Skottsberg formally published the Donatiaceae. The APG II system recommended the inclusion of "Donatia" in Stylidiaceae but allowed for the optional recognition of the family Donatiaceae. The APG III system merges Donatiaceae into Stylidiaceae. The two species in the genus represent a wide geographic range. "D. novae-zelandiae" is found in the alpine and subalpine regions of New Zealand and Tasmania while "D. fascicularis" is native to similar habitats in southern South America to latitude 40°S.
In Chile "Donatia fascicularis" is, together with "Astelia pumila", dominant in the cushion bogs that exists in areas exposed to the Pacific coast. As such it is not usually found together with "Sphagnum" which tend to grow slightly more inland.