Amaranthoideae

The Amaranthoideae are a subfamily of the Amaranthaceae. The stamens have anthers with two lobes (locules) and four pollen sacs. 
The main distribution of the subfamily is in tropical America, in tropical and Southern Africa, and in Australia.
The genera "Amaranthus" (the amaranths) and "Celosia" (the cockscombs) contain many ornamental species, as well as species whose seeds are used as pseudocereals and leaves as leaf vegetables.
Systematics.
The subfamily Amaranthoideae comprises about 57 genera with about 330 species. Phylogenetical research revealed that the subfamily is polyphyletic and its traditional classification (tribe Amarantheae Rchb. with two subtribes Amaranthinae und Aervinae) does not reflect the phylogenetic relationship. Therefore, a new taxonomical grouping is required. Müller & Borsch (2005) recognized several clades: