Eonatator

Eonatator is an extinct genus of marine lizard belonging to the mosasaur family. It is a close relative of "Halisaurus", and part of the same subfamily, the Halisaurinae. It is known from the Late Cretaceous of North America, Colombia and Sweden. Originally, this taxon was included within "Halisaurus", but was placed in its own genus, which also led to the subfamily Halisaurinae being created for the two genera.
Discovery and naming.
"Eonatator" is known from the Smoky Hill Chalk Member of the Niobrara Chalk Formation (Late Coniacian to Early Campanian) of Kansas, from the Eutaw Formation (Santonian) and Mooreville Chalk Formation (Selma Group; Santonian-Lower Campanian) of Alabama (United States), from the Kristianstad Basin of southern Sweden (late early Campanian), and the unit Nivel de Lutitas y Arenas (Campanian) of the Olini Group in La Mesa, Colombia.
The name "Eonatator" means "dawn swimmer" (Greek "eos" = dawn + Latin "natator" = swimmer). Originally, it contained only a single species, "E. sternbergii". The species is named in honour of Charles H. Sternberg and his son, Levi, who discovered the type specimen in the Niobrara Chalk during the summer of 1918. A second species, "E. coellensis", was named for the town of Coello in the Department of Tolima in Colombia, near of which it was discovered.
Description.
"Eonatator" was among the smaller mosasaurs. The length of the type skeleton, which represents an adult, is only 2.65 meters in length. Like many mosasaurs, it likely fed primarily on fish and smaller marine reptiles. The length of the type specimen of "E. coellensis", IGM p 881237 (which lack a complete tail), is of 2.8 meters until the last preserved caudal vertebrae, with a skull of 41,5 centimeters. The specimen of this species is remarkable for having remains of soft tissue in the ear region, the neck, thoracic and the abdominal region. Under the pygal vertebrae and the seventeenth dorsal vertebra there is a series of 20 small vertebrae centra and a flattened bone, that together measure 25 centimeters in length. It have features of the mosasauroids, with three vertebrae with haemal arches and procoelic centra, that suggest the possibility that these small bones belong to an embryo of this species, although the lack of diagnostic fossils like the skull or teeth prevents a complete identification. In any case, it will be consequent with the ovoviviparism previously reported in mosasauroids like "Carsosaurus".
Bardet et al. (2005, p. 465) diagnose "Eonatator sternbergii" as follows: "Ambiguous characters: premaxilla-maxilla lateral suture ending posterior to 9th maxillary teeth; tail about 40% of the head and trunk length (convergent in mosasaurines); caudal vertebra length greater than width; fewer than four pygal vertebrae; femur length about twice distal width (convergent in "Clidastes"). Autapomorphies: parietal with smooth triangular table extending very far posteriorly, bearing medium-sized circular foramen, located at distance twice its diameter from the frontal-parietal suture, and surrounded anteriorly and posteriorly by two parallel ridges; rounded quadrate with regularly convex tympanic "ala" (wing); vertebral formula: seven cervicals, 24 dorsals, four pygals, 28 median caudals and at least 41 terminal caudals; humerus length approximately 2.5x distal width." "E. coellensis" is diagnosed by more retracted nostrils, between the 7 and the 17 maxillary teeth, premaxilla and maxilla with a short rostrum anterior to the first teeth; presence of a septomaxilla, a large prefrontal that makes most of the margin of the outer nostril, a short and wide frontal, a parietal foramen located near of the fronto-parietal suture, a triangular surface of the parietal with two medial depressions and 22 caudal vertebrae.
Classification.
Like many mosasaurs, this genus has a complicated taxonomic history. The type specimen (UPI R 163, Uppsala University Palaeontological Institute, Uppsala, Sweden), a nearly complete skeleton, was originally referred to the genus "Clidastes" by Wiman and then to "Halisaurus" by Russell. Hence, "Clidastes sternbergii" became "Halisaurus sternbergii". However, by the late 1980s, some paleontologists began to suggest that "H. sternbergii" belonged in its own genus and that "Halisaurus" was polyphyletic.
In 2005, "Halisaurus sternbergii" was reassigned to its own genus, "Eonatator" by Nathalie Bardet and colleagues along with the description of "Halisaurus arambourgi" and the creation of the subfamily Halisaurinae.
Below is a cladogram following an analysis by Takuya Konishi and colleagues (2015) done during the description of "Phosphorosaurus ponpetelegans", which showcases the internal relationships within the Halisaurinae. The analysis excluded the dubious "Halisaurus onchognathus" and the genus "Pluridens".