TWENTY FIVE NEW SPECIES OF CISSEIS (SENSU STRICTO) AND TWO NEW SYNONYMS (COLEOPTERA: BUPRESTIDAE: AGRILINAE)

Summary Twenty five new species of Cisseis (sensu stricto) are described: Cisseis aenea; C. aenigma; C. aeruginosa; C. bilyi; C. borealis; C. chlorota; C. cupreata; C. erythrocephala; C. fritilla; C. furfurosa; C. goldingi; C. guttata; C. leai; C. minuta; C. miyama; C. montana; C. rubricata; C. rubriceps; C. storeyi; C. suehasenpuschae; C. tyleri; C. ustulata; C. wagneri; C. williamsi; C. wilsonensis. Two new synonyms of Cisseis (s.s.) are proposed: C. septuosa Barker 2001 = C. scabiosa (Boisduval, 1835) and C. spilota (Hope, 1846) = Pachycisseis bicolor (Gory & Laporte, 1839).


Introduction
The genus Cisseis Gory & Laporte (1839) has not been revised since Carter (1923) gave a synonymy and described new species. Since then many more specimens have become available, mainly through the activities of amateur collectors. Much of this material has been acquired by ANIC and is available for scientific study. Since 1997 I have been working towards a complete revision of the genus and for this purpose have examined all of the readily available type material. To date I have published a number of redescriptions, selected lectotypes, described twenty three new species and presented a check list of species and the location of type material (Barker 1998(Barker , 1999a(Barker , 1999b(Barker , 2001(Barker , 2002. Together with Dr C. L. Bellamy I have reassigned a number of species placed in Cisseis to a new genus Stanwatkinsius (Barker & Bellamy, 2001). In this paper I describe twenty five new species and recognise two synonyms.
Shape & sculpture: body dorso-ventrally flattened. Head punctured, shallow median sulcus, interantennal bridge 0.15 inter-ocular width. Pronotum striolate, apical margin projecting broadly medially, basal margin sinuate, large shallow fovea at each angle close to base, dorsal carina diverging from ventral carina basally and continuing until meeting anterior margin. Scutellum scutiform, unpunctured, anterior margin convex. Elytra scutellate, laterally more or less parallel-sided from base, rounded from post-medial to apex, with clumped setae forming the following faint patterns on each elytron, thick band at middle, four clumps across pre-apical area. Ventral surface faintly striolate laterally, glabrous medially, with sparse setae except laterally on abdominal segments clumps getting progressively smaller posteriorly. Legs: tarsal claws without inner tooth.
Colour: head coppery. Antennae bronze with coppery reflections. Rest of body bronze.
Shape & sculpture: as in male.

Remarks
This species could be confused with Cisseis patricia Carter, described from WA but also occurring in NSW. It is a slightly smaller species than C. patricia, the dorsal surface is highly reflective in C. aenea while it is dull in C. patricia. Males of C. patricia do not have any green colouration. Male genitalia differ from those in C. aenea.

Etymology
The species is named for its colour aeneus L., brazen (of copper).
Colour: head green with yellow reflections. Antennae green. Pronotum and scutellum green with yellow reflections. Elytra black with white markings formed from clumped setae. Ventral surface and legs green.
Shape & sculpture: head punctured, median sulcus, inter-antennal bridge 0.15 inter-ocular width. Pronotum striolate, anterior margin projecting medially, basal margin sinuate, dorsal carina diverging from ventral carina basally, then more or less parallel-sided until meeting anterior margin, clumped setae forming a white band on each side from post-medial to pre-medial near dorsal carina, space between carinae filled with setae. Scutellum scutiform, punctured, anterior margin convex, sides prolonged laterally. Elytra scutellate, laterally angled out slightly from base, rounded at humeral callus, more or less parallel-sided until rounded postmedially and tapered to pre-apex, rounded at apex, apical margin sub-serrate, with the following white spots on each elytron: two at base, one closer to the margin than middle, the other closer to middle than margin; one close to margin at humeral callus; one posterior premedial closer to suture than margin; one medial close to margin; one post-medial close to margin; one pre-apical closer to suture than margin; the last four forming a semi-circle in the middle of which are several marks close to the suture. Ventral surface striolate, setae moderately long, laterally clumped at meta-coxae and on visible abdominal segments to form white spots. Legs: tarsal claws with inner tooth.
Shape & sculpture: as in male.

Remarks
Male genitalia of this species resemble those in Cisseis vicina Kerremans (Barker 2001, Fig . 1K). C. vicina is a much smaller species and it lacks setae on the pronotum.

Etymology
The species remained unrecognised for more than fifty years and is named aenigma L. mystery. Colour: all dark green.

Colour: as in male.
Shape & sculpture: as in male.

Remarks
In shape and texture this species is closest to C. roseocuprea (Hope). That species however is all bronze, the tarsal claws are without inner teeth and the male genitalia differ (Barker 2001, Fig. 1E).
There is no definitive locality for the two specimens available from NSW.

Etymology
The species is named for its colour aeruginosus L. greenish-blue. Colour: head green. Antennae bronze. Pronotum and scutellum green. Elytra dark purple with coppery basal margin and base of suture. Ventral surface and legs green. Setae silver.
Shape and sculpture: head sparsely punctured, with faint median sulcus, inter-antennal bridge 0.23 inter-ocular width. Pronotum punctate-striolate, apical margin projecting broadly, basal margin sinuate, dorsal carina diverging from ventral carina at base and then more or less straight to apical margin. Scutellum scutiform, shallowly punctured. Elytra scutellate, laterally faintly rounded at humeral callus then almost parallel-sided until post-medially rounded and tapered, rounded at apex, apical margin sub-serrate, with clumps of setae forming irregular markings mainly in pre-and post-medial area with a single round pre-apical mark in the middle on each elytron. Ventral surface faintly scutellate with short setae, except clumps of pubescent setae on lateral coxae and abdominal sternites. Legs: tarsal claws with small inner tooth.
Colour: head, pronotum and scutellum blue-green, rest as in male.
Shape & sculpture: as in male.

Remarks
This species is closest to C. rubicunda Kerremans and about the same size. In C. rubicunda both sexes have a coppery head, pronotum and ventral surface; the elytra are coppery basally and mottled purple apically; male genitalia are different although of the same general form.

Etymology
The species is named to honour Dr S. Bílý, Coleopterist at NMPC. Colour: head variably green, coppery at base. Antennae, antennomeres: 1-2 coppery; 3-11 yellow bronze. Pronotum and scutellum coppery. Elytra black with dark blue reflections and clumps of setae forming white spots. Ventral surface and legs yellow bronze.

Cisseis borealis
Shape & sculpture: head flat, inter-antennal bridge 0.17 inter-ocular width. Pronotum striolate, apical margin projecting medially, basal margin sinuate, dorsal carina diverging from ventral carina at basal margin, more or less parallel then diverging before reaching apical margin. Scutellum scutiform, without punctures, medially excavated, anterior margin convex. Elytra scutellate, with short setae beneath each plate pointing posteriorly, apical margin sub-serrate, with following white markings on each elytron formed from clumps of pubescent setae: one in middle at base; one beneath humeral callus; four arranged in a semi-circle from pre-medial to pre-apical areas, anterior closer to suture than margin, two closer to margin than suture, most posterior largest, closer to suture than margin. Ventral surface striolate, sternum with deep punctures lying in striae, shallowly on abdomen with short setae projecting posteriorly except clumps of pubescent setae laterally on meta-sternal coxae and abdominal segments becoming progressively smaller, absent on last segment. Legs: tarsal claws with small inner tooth.
Colour: head coppery, rest as in male.
Shape & sculpture: as in male.

Remarks
C. rubriceps sp. nov. has a similar but not identical colour combination. The structure of the male genitalia readily separates them (Fig. 1j).

Etymology
The species is named for its geographic occurrence, borealis L. northern.
Colour: head green, mostly with yellow reflections, coppery on inter-antennal bridge. Antennae green, antennomeres 1-2 with coppery reflections. Pronotum green with yellow reflections medially, coppery laterally. Scutellum green with coppery reflections. Elytra mainly green, yellow green along suture. Ventral surface and legs coppery.
Colour: as in male.
Shape & sculpture: as in male.

Remarks
This species is unlike typical Cisseis species. The whole dorsal surface is densely punctured; the dorsal carina on the pronotum is indistinct; there are no clumps of setae forming marks or patterns on the elytra; there is no tibial setigeris on the legs; the median sulcus gives it the superficial appearance of members of the genus Ethonion.

Etymology
The species is named for its colour chloros Gr. green. Colour: head mostly green, bronze at base. Antennae black with bronze reflections. Pronotum bronze-green, green or coppery. Scutellum black with bronze reflections. Elytra black with purple or blue reflections and white marks formed from clumped setae. Ventral surface and legs black with bronze reflections.

Cisseis cupreata
Shape & sculpture: head punctate, flat, inter-antennal bridge 0.2 inter-ocular width. Pronotum striolate, anterior margin projecting medially, basal margin sinuate, laterally bulbous, space between the two carinae visible from above for half its length from base, dorsal carina diverging from ventral carina at base, more or less parallel-sided until approaching but not meeting anterior margin. Scutellum scutiform, flat, without punctures, sides extended laterally and rounded, anterior margin convex. Elytra scutellate, laterally more or less parallel-sided from base, rounded and tapered post-medially, rounded at apex, with the following white marks formed from clumped setae: four on each elytron forming a semi-circular pattern, first pre-medial closer to suture than margin, the second and third medial and post-medial closer to margin than suture, the fourth pre-apical closer to suture than margin, several smaller marks in middle along the suture. Ventral surface scutellate with short setae. Legs: tarsal claws with very small inner tooth; meta-tibial setigeris with two raised clumps.
Colour: head coppery. Antennae black with bronze reflections. Pronotum coppery. Scutellum black. Elytra black with markings as in male. Ventral surface and legs as in male.
Shape & sculpture: as in male.

Remarks
This species could be confused with Cisseis cupreola Barker which is a similar colour to males of this species. However, the colour combination of the sexes, the bulbous shape of the pronotum and the structure of the male genitalia (Barker 2001, Fig. 1P) separate them.

Etymology
The species is named for its colour cuprum L., copper. Colour: head red with blue reflections. Antennae, antennules: 1-6 coppery; 7-11 bronze. Pronotum and scutellum bronze. Elytra black with blue and purple reflections and with white spots. Ventral surface and legs coppery.
Shape & sculpture: head closely punctured with an impressed line at base, inter-antennal bridge 0.25 inter-ocular width. Pronotum striolate, anterior margin projecting medially, posterior margin sinuate, laterally parallel-sided from base rounded pre-medially to anterior margin, dorsal carina diverging from ventral carina basally then more or less parallel-sided until ending just before the anterior margin. Scutellum scutiform, with few punctures, anterior margin convex. Elytra scutellate, laterally more or less parallel-sided from base, rounded postmedially and tapered to rounded, sub-serrate apices. Ventral surface faintly striolate with sparse, short setae except laterally on meso-and meta-coxae and abdominal segments, dense clumps of setae form white spots decreasing in size posteriorly. Legs: tarsal claws without inner tooth.
Colour: head coppery, rest as in male.
Shape & sculpture: as in male.

Remarks
This species has the same coloration as Cisseis cupreola Barker except that the male head differs. Cisseis erythrocephala sp. nov. can be distinguished by being smaller, elongate and the male genitalia differ (Barker 2001, Fig. 1P).

Etymology
The species is named for the head colour of the male erythros Gr. red, kephale Gr. head.
Colour: head green with purple reflections. Antennae purple. Pronotum green with coppery reflections, laterally coppery. Scutellum green. Elytra dark red with purple reflections and white marks formed from clumped setae. Ventral surface and legs coppery. Setae silver.
Shape & sculpture: head shallowly punctured, faint median sulcus, inter-antennal bridge 0.2 interocular width. Pronotum shallowly striolate, anterior margin projecting medially, posterior margin sinuate, laterally dorsal carina angled away from ventral carina at base then running more or less parallel to it until just after the middle where it ends, wide interval between carina densely covered with setae. Scutellum scutiform, with few punctures, anterior margin convex. Elytra scutellate, laterally angled out from base, rounded at humeral callus, more or less parallel-sided, rounded after middle then tapered, rounded at apex, apical margins serrate, with following prominent white spots on each elytron formed from clumped setae: one in middle near anterior margin, one directly posterior to it, four in semicircle, first pre-medial closer to suture than margin, second pre-medial closer to margin than suture, third post-medial closer to margin than suture, fourth pre-apical closer to suture than margin, with smaller irregular spotting around middle close to suture and one small spot behind humeral callus close to margin, absent in holotype but present in two other specimens. Ventral surface scutellate with short setae, laterally with white spots formed from clumped setae on metacoxae and abdominal segments. Legs: tarsal claws with small inner tooth.
Colour: as in male, except that the elytra are brighter red.
Shape & sculpture: as in male.

Remarks
This is one of a number of species with heavily spotted elytra. The colour is similar to that of male C. derbyensis Barker, which is larger with different male genitalia (Barker 2001, Fig. 1AB).

Etymology
The species is named for its prominent spotting fritillus L., spotted. Colour: head mostly bronze, coppery basally. Antennae bronze. Pronotum and scutellum coppery. Elytra brown with coppery reflections basally. Ventral surface dark with coppery reflections. Legs dark with coppery and green reflections.
Shape & sculpture: head closely punctured, with median fovea; inter-antennal bridge 0.28 interocular width. Pronotum striolate, broadly projecting medially, dorsal carina diverging from ventral carina at base, then more or less parallel-sided, diverging just before reaching anterior margin. Scutellum scutiform, with few punctures, expanded laterally, anterior margin convex. Elytra scutellate, with clumps of short setae not defined into definite patterns except in the pre-apical area where they form a broad fascia from margin to suture and a posterior clump, apical margin rounded and sub-serrate. Ventral surface faintly scutellate moderately setose, more so laterally than medially. Legs: tarsal claws with prominent inner tooth.
Colour: head coppery. Antennae bronze. Pronotum and scutellum coppery. Elytra dark with coppery reflections. Ventral surface and legs dark with coppery reflections.
Shape and structure: as in male.

Remarks
This species is closest to Cisseis trimentula Barker which also has accessory structures on the parameres (Barker 2001, Fig. 1G). That species is much smaller and darker and occurs only in NSW. Either Jan Obenberger or one of his collectors mis-named many specimens from Cooktown, Queensland 'Yorktown.' This was commented on by Carter (1940) who stated incorrectly that a Yorktown didn't exist, thus ignoring Yorketown, South Australia.

Etymology
The name is derived from furfurosus L., brownish.
Colour: head coppery-red. Pronotum blue-green with coppery-red reflections. Scutellum and elytra dark blue-green with coppery-red reflections. Ventral surface and legs coppery-red.
Shape & sculpture: head closely punctured, flat, inter-antennal bridge 0.2 inter-ocular width. Pronotum striolate, anterior margin slightly projecting medially, basal margin sinuate, dorsal carina diverging from ventral carina basally then more or less parallel to it, converging slightly then diverging to meet anterior margin. Scutellum scutiform, few punctures, anterior margin convex. Elytra scutellate, laterally tapering post-medially to rounded sub-serrate apex, with the following very faint white marks formed from small clumps of setae: circle formed from four spots on each elytron around middle, one pre-medial closer to suture than margin, second medial closer to margin than suture, third post-medial closer to margin than suture, fourth pre-apical closer to suture than margin, fifth in centre close to suture. Ventral surface faintly scutellate, with lateral white spots formed from clumped setae on each segment commencing with largest on meta-sternum, becoming progressively smaller along abdomen. Legs: tarsal claws without inner tooth.

Remarks
This species is distinct from all others found in WA being small and green and with distinctive male genitalia.
Colour: head, antennae, pronotum and scutellum bronze. Elytra black with white spots. Ventral surface and legs green with bronze reflections.
Shape & sculpture: head with median sulcus, inter-antennal bridge 0.16 inter-ocular width. Pronotum striolate, anterior margin projecting medially, basal fovea with pubescent setae on each side, dorsal carina diverging from ventral carina basally then more or less parallel-sided reaching anterior margin, with two patches of pubescent setae between carina and two matching patches on the dorsal side of the dorsal carina. Scutellum extended laterally at convex anterior margin, punctured. Elytra scutellate, apically sub-serrate, with the following white marks on each elytron: two at base, one beneath the humeral callus, four forming a semicircle from pre-to post-medial, the first and the last closer to the suture than margin, the second and third closer to the margin than suture, the third elongate the rest more or less circular, with several small obscure marks in the centre close to the suture. Ventral surface faintly scutellate, with short setae except laterally with clumped setae on meta-tibial coxae and abdominal segments becoming progressively smaller posteriorly. Legs: tarsal claws with inner tooth.
Shape & sculpture: as in male.

Remarks
This species is distinguished by the dichromatic colour and very distinct spotting on the upper surface. It is unlike any other known W.A. species.

Etymology
The species is named for its elytral marking guttata L., spotted.
( Fig. 1i,  Colour: head dark blue with bronze and coppery reflections. Pronotum bronze with coppery and purple reflections. Scutellum, elytra, ventral surface and legs black with blue and coppery reflections.
Colour: as in male.
Colour: head mainly green, bronze and coppery at base. Antennae black with coppery reflections. Pronotum brown medially with bronze reflections laterally. Scutellum brown with bronze reflections. Elytra brown with bronze reflections and faint white marks formed from clumped setae. Ventral surface and legs brown with bronze reflections.
Shape & sculpture: head punctured, with shallow median sulcus, inter-antennal bridge 0.2 interocular width. Pronotum striolate, anterior margin projecting medially, basal margin sinuate, laterally dorsal carina diverging from ventral carina at base then gradually diverging until it reaches anterior margin. Scutellum scutiform, with few punctures, anterior margin convex. Elytra scutellate with moderately long setae scattered over basal half except for two glabrous areas in pre-medial area, post-medial zig-zag fascia formed from clumped setae. Ventral surface with shallow punctures, sparse short setae. Legs: tarsal claws with small notch.
Colour: head coppery-red, rest as in male.
Shape & sculpture: as in male.

Remarks
This is one of the smallest species of Cisseis to be recognised so far. The colour combination does not resemble any of the other small species known.

Etymology
The species is named for its small size minutus L., small.
Colour: head mostly green, bronze at base. Antennae bronze. Pronotum background colour black, striae coppery. Scutellum bronze. Elytra black with setae forming white patterns and marks. Ventral surface: prosternal process green; dorsal surface of first femora green; remaining legs and ventral surface coppery.
Shape and sculpture: head punctured, prominent median sulcus, inter-antennal bridge 0.14 interocular width. Pronotum prominently striolate, anterior margin projecting broadly, posterior margin sinuate, dorsal carina diverging from ventral carina basally then more or less parallel to it, diverging towards the anterior margin but not reaching it. Scutellum scutiform, punctured, anterior margin convex. Elytra basal half striolate, apical half scutellate, with clumps of white setae forming a variable pattern across the middle and a wavy line across the pre-apical area, apex rounded and sub-serrate. Ventral surface laterally faintly scutellate, medially glabrous, with sparse short setae adpressed to plates and pointing posteriorly. Legs: tarsal claws with prominent medial tooth.
Colour: head mainly coppery-red, bronze at base, labrum green. Rest of upper surface as in male. Ventral surface black, legs black with faint purple reflections.
Shape and sculpture: as in male.

Remarks
This species can be distinguished by the male genitalia. It does not resemble any of the described species from Australia.

Etymology
The species name is derived from the type locality. Colour: head green. Antennae bronze with coppery reflections. Pronotum, scutellum, elytra, ventral surface and legs dark brown with bronze and coppery reflections.
Shape and sculpture: cylindrical. Head punctured, flat, inter-antennal bridge 0.12 inter-ocular width. Pronotum striolate, anterior margin projecting medially, posterior margin sinuate, laterally widest near base then tapering to anterior margin, dorsal carina gradually diverging from ventral carina basally until it meets anterior margin. Scutellum scutiform, with few punctures. Elytra scutellate, laterally more or less parallel-sided from base, rounded postmedially then tapered until rounded at apex, with faint, irregular spotting medially formed from clumps of setae and two on each elytron in pre-apical area, first closer to margin than suture, second posterior, closer to suture than margin. Ventral surface faintly scutellate with sparse, short setae. Legs: tarsal claws without inner tooth; metatibial setigeris from middle to distal end.
Colour: head coppery. Rest as in male.
Shape & sculpture: as in male.

Remarks
This species could be confused with other small brown species particularly C. patricia Carter. However, C. patricia is dorsoventrally flattened while C. montana is cylindrical and the male genitalia differ.

Etymology
The name of this species refers to its distribution in the Blue Mountains and is derived from montanus L., of mountains. Colour: head, antennae, pronotum and scutellum coppery. Elytra coppery at base merging into black with coppery reflections pre-medially. Ventral surface: pro-sternum coppery, meso-and meta-sternum bronze, abdomen green with bronze reflections. Legs bronze. Setae silver.
Shape & sculpture: head closely punctured, no median sulcus, inter-antennal bridge 0.13 interocular width. Pronotum striolate, anterior margin projecting medially, basal margin sinuate, laterally dorsal carina diverging from ventral carina at base, then more or less parallel-sided not quite reaching the anterior margin. Scutellum scutiform, punctured anterior margin barely convex. Elytra scutellate, laterally more or less parallel-sided from base, rounded postmedially and tapered, rounded at apex, apical margin sub-serrate, with following white spots formed from clumped setae: one obscure medially at anterior margin; one obscure pre-medial closer to suture than margin; one distinct pre-medial closer to margin than suture; one distinct post-medial closer to margin than suture; one obscure pre-apical closer to suture than margin. Ventral surface shallowly punctured, short sparse setae. Legs: tarsal claws without inner tooth.
Colour: dorsal surface as in male. Ventral surface and legs coppery.
Shape & sculpture: as in male.

Remarks
This species could be confused with C. rubriceps sp. nov. However, the elytral markings are less prominent and male genitalia are very different (Fig. 1j).

Etymology
The species is named for its colour rubicundus L., ruddy.
Shape & sculpture: head with median impressed line from middle to base, inter-antennal bridge 0.2 inter-ocular width. Pronotum striolate, apical margin projecting medially, dorsal carina diverging from ventral carina at base then more or less parallel to it until diverging just before reaching apical margin, white setae along dorsal edge of dorsal carina and in clump between carinae at apical margin. Scutellum scutiform, extending laterally at convex basal margin, punctured. Elytra with apical margin sub-serrate, with following white spots on each elytron formed by clumps of setae: two along anterior margin; four forming semicircle comprised of large round anterior spot in middle of pre-medial area close to suture, two smaller spots closer to margin than suture, one medial other post-medial, one elongate in pre-apical area closer to suture than margin; two smaller spots close to suture, one behind other in medial area which may have small accessory spots. Ventral surface faintly scutellate with very short setae except dense clumps of pubescent setae at lateral margins, largest on metathoracic sternite, becoming progressively smaller on abdominal sternites. Legs: short metatibial setigeris from post-medial to distal end; tarsal claws with no inner tooth.
Colour: head coppery, rest as in male. One female specimen has green head and pronotum.
Shape & sculpture: as in male.

Remarks
The Mount Barker specimens were collected on the flowers of Eucalyptus sp. Of the specimens examined, females had more conspicuous spotting on the elytra than males. Cisseis rubricata sp.
nov. and Cisseis modesta Kerremans are both small red species also found in W.A., both are smaller than Cisseis rubriceps sp. nov. and male genitalia differs (Fig. 1t).
Shape & sculpture: head very closely and deeply punctured with median sulcus, inter-antennal bridge 0.14 inter-ocular width. Pronotum deeply striolate, dorsal carina more or less straight from base, diverging from ventral carina for its whole length but not quite reaching anterior margin. Scutellum scutiform, punctured, sides extended laterally and rounded to meet anterior margin. Elytra heavily scutellate, laterally angled out from basal margin, rounded at humeral callus, more or less parallel-sided until post-medially rounded and narrowed to rounded apex, apical margins sub-serrate, with the following white markings on each elytron formed from clumped setae: one at base closer to margin than middle, one at humeral callus, four in semicircle the first pre-medially close to suture, the second posterior to first and closer to margin, the third post-medial and posterior to second close to margin, the fourth pre-apical and close to suture, one in middle close to suture. Ventral surface heavily striolate, moderately setose, setae moderately long, laterally clumped to form white marks on meta-coxae and all visible abdominal segments. Legs: meta-tibial setigeris raised in two places; tarsal claws with small inner tooth.

Remarks
This species could be confused with Cisseis borealis sp. nov. but it is larger and male genitalia are quite different (Fig. 1d).
Shape & sculpture: as in male.

Remarks
There is no other described species of Cisseis which is this shape, reminiscent of Stanwatkinsius (Barker & Bellamy, 2001) and without dorsal markings. Male genitalia resemble C. albosparsa Gory & Laporte, but that species has elytral spotting and is all coppery on the upper surface, except green head of male.

Etymology
The species is named after Sue Hasenpusch, 'Garradunga', Innisfail. Colour: Head mainly green, brown at base, rest of body brown.
Shape & sculpture: head closely punctured, inter-antennal bridge 0.18 inter-ocular width. Pronotum striolate, apical margin projecting medially, basal margin sinuate, laterally dorsal carina angled away from ventral carina at base then more or less parallel-sided until joining anterior margin. Scutellum scutiform, few punctures, anterior margin convex, elongate at sides. Elytra scutellate, laterally angled out from base, rounded at humeral callus, more or less parallelsided until rounded after middle, tapered then rounded at apex, apical margins sub-serrate. Ventral surface with shallow punctures, short sparse setae. Legs: tarsal claws without an inner tooth.
Colour: head mainly coppery, brown at base, rest brown.
Shape & sculpture: as in male.

Remarks
This species is unique and very easy to identify because of the colour difference between the sexes, size difference and male genitalia.
Colour: head mainly bright green, bronze basally with coppery reflections. Antennae, antennomeres: 1-4 bronze with green reflections; 5-11 bronze. Pronotum black, bottoms of striae with coppery reflections; space between carinae coppery with green and bronze reflections. Scutellum bronze. Elytra black with coppery and bronze reflections and clumps of silver setae forming patterns. Ventral surface black, sternal segments with green reflections medially. Legs: 1 & 2 with dorsal surface of femora green, ventral surface black; 3 all black.
Shape & sculpture: head shallowly punctured from apex to middle then shallowly striolate to base, broad medial sulcus, inter-antennal bridge 0.22 inter-ocular width. Antennae, antennomeres: 1-3 obconic; 4-11 triangular. Pronotum striolate, apical margin projecting broadly medially, basal margin sinuate, with a broad basal fovea on each side, laterally dorsal carina diverging from base then more or less parallel to ventral carina until diverging just before it ends, not reaching apical margin. Scutellum scutellate, punctured, basal margin expanded laterally. Elytra scutiform, laterally more or less parallel-sided from base, rounded post-medially and tapered, rounded at apex, apical margin sub-serrate, with the following white pattern formed from clumps of setae: irregular marks from pre-to post-medially; pre-apical zig-zag fascia. Ventral surface: sternal segments striolate; abdominal segments shallowly punctate, whole surface with sparse, very short setae. Legs: tarsal claws with small inner tooth.
Colour: head mainly red, bronze basally. Pronotum as in male. Scutellum and elytra bronze. Ventral surface and legs black.
Shape & sculpture: as in male.

Remarks
This species resembles C. miyama sp. nov. but male genitalia are different (Fig. 1s).

Etymology
The species is named to honour the collector, the late Rev. L. Wagner. Colour: head green, black basally. Antennae green. Pronotum black with bronze reflections except ventral carina green. Scutellum bronze with green reflections. Elytra black with bronze reflections pre-medially and clumps of setae forming irregular white patterns. Ventral surface: prosternum green, rest bronze. Legs: dorsal femora green ventral femora and tibiae bronze. Setae silver.

Cisseis williamsi
Shape & sculpture: head closely punctured, deep median sulcus with a medial indentation on each side, inter-antennal bridge 0.25 inter-ocular width. Pronotum striolate with a large fovea on each side closer to basal margin than anterior margin, anterior margin projecting medially, basal margin sinuate, laterally dorsal carina angled away from ventral carina at base, more or less parallel-sided until reaching anterior margin. Scutellum scutiform, punctured, excavate. Elytra scutellate, laterally angled out from base, rounded at humeral callus and then more or less parallel-sided until rounded post-medially and tapered to rounded apices, apical margin sub-serrate, with the following markings formed from clumped white setae: diffuse marks from pre-medial to post-medial, broad post-medial fascia from margin to suture, narrow preapical fascia from margin to suture. Ventral surface with shallow punctures and short, sparse setae. Legs: tarsal claws with small inner tooth.
Colour: head coppery. Antennae black with coppery reflections. Pronotum and scutellum coppery. Elytra black with white markings. Ventral surface black. Legs black with coppery reflections on dorsal surfaces.
Shape & sculpture: as in male.

Remarks
This species could be confused with two others namely C. trimentula Barker and C. watkinsi Barker which are about the same size and colour except for the colour of the head, black in C. trimentula and dull green or blue in C. watkinsi. The structure of the male genitalia is different in all three species (Barker 2001, Fig.1).

Etymology
Named after the collector G. Williams, Lansdowne.
Colour: head green. Antennae yellow-green. Pronotum green with yellow reflections. Scutellum green. Elytra black with white markings formed from clumped setae. Ventral surface: sternites and fused abdominal segments green with yellow reflections; last three abdominal segments black with yellow reflections. Legs green with yellow reflections.
Shape & sculpture: head punctured, median sulcus, inter-antennal bridge 0.27 inter-ocular width. Pronotum striolate, apical margin broadly projecting medially, basal margin sinuate, dorsal carina more or less straight from base and gradually diverging from ventral carina, meeting apical margin. Scutellum scutiform, without punctures, sides elongate along basal margin. Elytra striolate, laterally more or less parallel-sided rounded post-medially, tapered towards then rounded at apex, apical margin sub-serrate, with a few small, irregular white marks. Ventral surface faintly striolate with sparse, short setae. Legs: tarsal claws with prominent inner tooth.
Colour: head, antennae and pronotum dark brown with coppery reflections. Elytra dark brown with bronze and coppery reflections. Ventral surface and legs dark brown.
Shape & sculpture: as in male.

Remarks
This species could be confused with Cisseis marmorata Gory & Laporte but they differ in the following. C. marmorata males have brown elytra while they are black in this species. The females of C. marmorata have a bulbous pronotum not found in the female of this species. Male genitalia differ.
Etymology Named for the type locality.