SRL for [annealed]: A series of steps catalyzed in part by the recombinase then leads to joining of the two helices by at least one Holliday junction , in which a segment of a single strand in each helix is [annealed] to the complementary strand in the other helix .
ARG-1: a segment of a single strand in each helix | ARG-5: to the complementary strand in the other helix | ARG-M-LOC: which