this work was supported by feder funds and fondo social europeo through grants from the junta de andaluca ( grants cvi-7335 to t.k . and cvi-7391 to m.eu ) and the spanish ministry for economy and competitiveness ( grants bio2010 - 16937 and bio2013 - 42297 to t.k ; grant bfu2010 - 17946 to m.eu ) .