In this regard, expression of VvMYB4-like and the gentian GtMYB1R1 and GtMYB1R9 share the down-regulation of the endogenous DFR and ANS (Nakatsuka et al. 2013), whereas with the overexpression of the strawberry FaMYB1, it is shared only the down-regulation of ANS/ LDOX (Aharoni et al. 2001).