significant direct and indirect shareholdings (including indirect shareholdings through pyramid structures and cross-shareholdings) within the meaning of Article 85 of Directive 2001/34/EC; ###>les participations significatives au capital, directes ou indirectes (par exemple, des participations indirectes au travers de structures pyramidales ou d'actionnariat croisé), au sens de l'article 85 de la directive 2001/34/CE;