The programme also covers chemical soil properties, such as pH, BS, cation exchange capacity, C/N and soil organic C. Tree crown defoliation integrates forest health status, which reacts to many biotic, abiotic and anthropogenic stresses. ###>La défoliation de la cime des arbres témoigne de l'état de la forêt, qui réagit à de nombreuses contraintes biotiques, abiotiques et anthropiques.