The Sclerodermatineae was first legitimately used by Manfred Binder and Andreas Bresinsky in 2002 based on molecular analyses of nuclear ribosomal large subunit ( 25S ) rRNA sequence from 60 species of Boletales . This research was an extension of Binder 's 1999 graduate work , in which he argued for the need to recognize the molecular differences of the sclerodermatoid fungi . Sclerodermatineae is one of six lineages of the Boletales recognized as a suborder ; the others are the Boletineae , Paxillineae , Suillineae , Tapinellineae , and Coniophorineae . Of the nine genera assigned to the Sclerodermatineae , three are hymenomycetes ( Boletinellus , Gyroporus , and Phlebopus ) , and six are gasteroid ( Astraeus , Calostoma , Diplocystis , Pisolithus , and Scleroderma ) . Since the suborder 's original description , there have been several phylogenetic studies investigating the Sclerodermatineae . Some studies have revealed the existence of numerous cryptic species and have contributed to taxonomic expansion of the group . The " core " Sclerodermatineae include the genera Astraeus , Calostoma , Scleroderma , Pisolithus , Diplocystis , Tremellogaster ( all gasteroid ) , and the boletoid genus Gyroporus ; Phlebopus and Boletinellus resolved as sister to this core group .