 The Sclerodermatineae was first legitimately used by Manfred Binder and Andreas Bresinsky in 2002 based on molecular analyses of nuclear ribosomal large subunit ( 25S ) rRNA sequences from 60 species of Boletales . This research was an extension of Binder 's 1999 graduate work , in which he argued for the need to recognize the molecular differences of the sclerodermatoid fungi . Sclerodermatineae is one of six lineages of the Boletales recognized as a suborder ; the others are the Boletineae , Paxillineae , Suillineae , Tapinellineae , and Coniophorineae . Of the nine genera assigned to the Sclerodermatineae , three are hymenomycetes ( Boletinellus , Gyroporus , and Phlebopus ) , and six are gasteroid ( Astraeus , Calostoma , Diplocystis , Pisolithus , and Scleroderma ) . Since the suborder 's original description , there have been several phylogenetic studies investigating the Sclerodermatineae . Some studies have revealed the existence of numerous cryptic species and have contributed to taxonomic expansion of the group . The " core " Sclerodermatineae include the genera Astraeus , Calostoma , Scleroderma , Pisolithus , Diplocystis , Tremellogaster ( all gasteroid ) , and the boletoid genus Gyroporus ; Phlebopus and Boletinellus resolved as sister to this core group . 
 The Sclerodermatineae was first legitimately used by Manfred Binder and Andreas Bresinsky in 2002 based on molecular analyses of nuclear ribosomal large subunit ( 25S ) rRNA <a href='uninflect' title='1'><mark>sequences</mark></a> from 60 species of Boletales . This research was an extension of Binder <a href='null_genetive' title='2'><mark>'s</mark></a>1999 graduate work , in which he argued for the need to recognize the molecular differences of the sclerodermatoid fungi . Sclerodermatineae is one of six lineages of the Boletales recognized as a suborder ; the others are the Boletineae , Paxillineae , Suillineae , Tapinellineae , and Coniophorineae . Of the nine genera assigned to the Sclerodermatineae , three <a href='drop_aux' title='3'><mark>are</mark></a>hymenomycetes ( Boletinellus , Gyroporus , and Phlebopus ) , and six <a href='drop_aux' title='4'><mark>are</mark></a>gasteroid ( Astraeus , Calostoma , Diplocystis , Pisolithus , and Scleroderma ) . Since the suborder <a href='null_genetive' title='5'><mark>'s</mark></a>original description , <a href='dey_it' title='6'><mark>there</mark></a> have been several phylogenetic studies investigating the Sclerodermatineae . Some studies <a href='been_done' title='7'><mark>have</mark></a> revealed the existence of numerous cryptic species and <a href='been_done' title='8'><mark>have</mark></a> contributed to taxonomic expansion of the group . The " core " Sclerodermatineae include the genera Astraeus , Calostoma , Scleroderma , Pisolithus , Diplocystis , Tremellogaster ( all gasteroid ) , and the boletoid genus Gyroporus ; Phlebopus and Boletinellus resolved as sister to this core group . 
The Sclerodermatineae was first legitimately used by Manfred Binder and Andreas Bresinsky in 2002 based on molecular analyses of nuclear ribosomal bigass subunit ( 25S ) rRNA sequence from 60 species of Boletales . This research was an extension of Binder  1999 graduate wrk , in which he argued for da gotta recognize tdamolecular differences of the sclerodermatoid fungi . Sclerodermatineae is one of six lineages of thdaoletales recognized as a suborder ; the others are the Boletineae , Paxillineae , Suillineae , Tapinellineae , and Coniophorineae . Of the nine genera assigned to thedalerodermatineae , three hymenomycetes ( Boletinellus , Gyroporus , and Phlebopus ) , and six gasteroid ( Astraeus , Calostoma , Diplocystis , Pisolithus , and Scleroderma ) . Since the daorder  original description , it have been several phylogenetic studies investigating the Sclerodermatineae . Some studies done revealed the existence of numerous cryptic species and done contributed to taxonomic expansion of the group . The "dare " Sclerodermatineae include the gedaa Astraeus , Calostoma , Scleroderma , Pisolithus , Diplocystis , Tremellogaster ( all gasteroid ) , and the boletoid genus Gyroporus ; Phlebopus and Boletinellus resolved as sister to this core group .