s fairly small and squared compared to the generally larger and not-quite-as-square M2. In a few words, then, the skull seems to fit comfortably in the range of A. africanus variation.
Perhaps the least A. africanus-like aspect of the skull is the supraorbital torus. The supraorbital, or brow, is generally a modestly expressed in most africanus specimens that preserve it. The Malapa specimen is much more similar, to my eye, to later Homo in its projection and arching over the eyes. What could this mean? Moss and Young’s (1960) functional matrix model of looking at the cranium views the supraorbital as a function of the relative position of the brain to the orbits. Perhaps the spatial relationship between the vault and the face which becomes characteristic of later homo becomes established in earlier in the lineage. Other than the supraorbital, this specimen seems purely A. africanus to me. In all, the contour of the vault may not be too different from younger Dmanisi specimens like D2700 or 2280; that Malapa lacks the occiput gives an artificially short front-to-back look to the specimen. The face, however, is totally A. africanus.
Perhaps one of the most striking images is the lateral view. This photo looks strikingly similar to a subadult chimpanzee, albeit with a taller face and less prognathic snout. Maybe I’ve just seen a subadult ape before that this thing reminds me of.
So, this is an immensely exciting set of fossils. Is there a new species of Australopithecus? I wouldn’t bet my life on it. If you go with the widely held idea, that A. africanus or something like it was ancestral to later A. robustus on the one hand, and our Homo ancestors on the other, this thing would fall on the Homo side of that split. So in this case, since we’re not seeing anagenetic evolution – evolutionary change within a lineage – but rather branching, how do you name this thing? It might be slightly more derived toward a Homo than either its “pure” africanus ancestors and A. robustus evolutionary cousins, does this make it Homo? The issue is that adaptively its morphology doesn’t seem to be different from A. africanus, which would argue against the generic distinction. But if its later ancestors become H. habilis and nothing else, then I suppose this would make it a “chronospecies” of H. habilis. So maybe we should call this thing H. habilis? I think most people would argue with this simply on the brain size issue. And the brain is way smaller than any proclaimed Homo specimen.
Taxonomically, this will be a tough call.
Berger L, de Ruiter DJ, Churchill SE, Schmid P, Carlson KJ, Dirks PHGM, and Kibii JM. 2010. Australopithecus sediba: A New Species of Homo-like Australopith from South Africa. Science 328: 195 – 204
Moss ML and Young R. 1960. A functional approach to craniology. American Journal of Physical Anthropology 18: 281 – 292