Q:

Number of Commits between two Commitishes

How can I find the number of commits between two commitishes in git?
Additionally, is there some way that I could do the same with any project on GitHub (using the UI, not the API)?

A:

Before I give you an answer, consider this commit graph:
        o -----------
       /             \
... - A - o - o - o - B
       \         /
        o ----- o

Each o represents a commit, as do A and B (they're just letters to let us talk about specific commits).  How many commits are there between commits A and B?
That said, in more linear cases, just use git rev-list --count A..B and then decide what you mean by "between" (does it include B and exclude A? that's how git rev-list --count will behave).  In branchy cases like this, you'll get all the commits down all the branches; add --first-parent, for instance, to follow just the "main line".
(You also mentioned "commitish", suggesting that we might have annotated tags.  That won't affect the output from git rev-list, which only counts specific commits.)

Edit: Since git rev-list --count A..B includes commit B (while omitting commit A), and you want to exclude both end-points, you need to subtract one.  In modern shells you can do this with shell arithmetic:
count=$(($(git rev-list --count A..B) - 1))

For instance:
$ x=$(($(git rev-list --count HEAD~3..HEAD) - 1))
$ echo $x
2

(this particular repo has a very linear graph structure, so there are no branches here and there are two commits "between" the tip and three-behind-the-tip).  Note, however, that this will produce -1 if A and B identify the same commit:
$ x=$(($(git rev-list --count HEAD..HEAD) - 1))
$ echo $x
-1

so you might want to check that first:
count=$(git rev-list --count $start..$end)
if [ $count -eq 0 ]; then
    ... possible error: start and end are the same commit ...
else
    count=$((count - 1))
fi

A:

$ git log 375a1..58b20 --pretty=oneline | wc -l

Specify your start commit followed by your end commit, and then count the lines.  That should be the count of commits between those two commit ranges.  Use the --pretty=online formatting so that each commit takes up a single line.
As for the GUI in GitHub, I don't know of a way to accomplish this same task.  But that should be trivial, as the above is the possible way to do it directly in Git Bash.

