Microswimmers, including flagellated bacteria such as E. coli [1]} and motile, single-celled eukaryotes such as Chlamydomonas [2]}, are common in biology and usually exist in complex fluid environments in nature.
Systematic studies on the dynamics of microswimmers that can help us to understand their complex behavior will thus allow us to further our understanding of basic biology and provide a guide for developing artificial micromachines. The latter could have great potential in various technological and biomedical applications [3]}, [4]}, such as targeted drug delivery [5]} or therapies using microrobots [6]}.
