where the stoichiometry matrix \(S\)  is determined by the network
structure [1]}. The relationship between the rate \(v_c\) 
of carbon fixation by Rubisco and the rate \(v_o\)  of the Rubisco
oxygenase reaction depends nonlinearly on the ratio of the local
oxygen and carbon dioxide concentrations (here expressed as equivalent
partial pressures),
\(\frac{v_o}{v_c} = \frac{1}{S_R} \frac{P_{O2}}{P_{CO2}}\) 
