Definition of GRAVAMEN: The burden or gist of a charge; the grievance or injury specially complained of.In English ecclesiastical law. A grievance complained of by the clergy befo The Law Dictionary Featuring Black's Law Dictionary Free Online Legal Dictionary 2nd Ed.
Phosphate and sugar make up the sides of a DNA ladder.
The rungs that are in the DNA ladder molecule are nucleotides. They  are adenine, guanine, thymine and cytosine. Deoxyribose and  phosphate make up the backbone of the molecâ¦ule.