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The proton motive force arises from a combination of the concentration gradient of protons and a voltage gradient across the membrane (the protons are positively charged, so an imbalance of protons means that the side of the membrane with more protons has a positive charge relative to the side with fewer protons).
The proton-proton collision is the primary operation method, but the LHC has also collided protons with lead nuclei for two months in 2013 and used leadâlead collisions for about one month each in 2010, 2011, and 2013 for other investigations.The LHC's computing grid was (and currently is) a world record holder.Test' collisions restarted in the upgraded collider on 5 April 2015, reaching 6.5 TeV per beam on 20 May 2015 (13 TeV total, the current world record for particle collisions). Its second research run commenced on schedule, on 3 June 2015.