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This super coiling accommodates the stress energy and the DNA becomes stable with 10.6 bp per coil. Generally the density of the super coils a DNA is one super coil per 200bp or per ~20 turns of the helix. Negative super coils help in stabilizing certain DNA structures i.e. Z-DNA, cruciform DNA, triplex DNA. It also helps in unwinding of the DNA during replication as replication bubble or during transcriptional initiation as transcriptional bubble.
The replication factor C, or RFC, is a five-subunit protein complex that is required for DNA replication. The subunits of this heteropentamer are named Rfc1, Rfc2, Rfc3, Rfc4, and Rfc5 in Saccharomyces cerevisiae. RFC is used in eukaryotic replication as a clamp loader, similar to the Î³ Complex in Escherichia coli.Its role as clamp loader involves catalysing the loading of PCNA onto DNA. It binds to the 3' end of the DNA and uses ATP to open the ring of PCNA so that it can encircle the DNA.he subunits of this heteropentamer are named Rfc1, Rfc2, Rfc3, Rfc4, and Rfc5 in Saccharomyces cerevisiae. RFC is used in eukaryotic replication as a clamp loader, similar to the Î³ Complex in Escherichia coli.