when do bryophytes (including mosses liverworts and hornworts) first appear in the fossil record
Timeline of plant evolution The evidence of plant evolution changes dramatically in the Ordovician with the first extensive appearance of spores in the fossil record (Cambrian spores have been found, also). The first terrestrial plants were probably in the form of tiny plants resembling liverworts when, around the Middle Ordovician, evidence for the beginning of the terrestrialization of the land is found in the form of tetrads of spores with resistant polymers in their outer walls. These early plants did not have conducting tissues, severely limiting their size. They were, in effect, tied to wet terrestrial environments by their inability to conduct water, like extant liverworts, hornworts, and mosses, although they reproduced with spores, important dispersal units that have hard protective outer coatings, allowing for their preservation in the fossil record, in addition to protecting the future offspring against the desiccating environment of life on land. With spores, plants on land could have sent out large numbers of spores that could grow into an adult plant when sufficient environmental moisture was present.
Radiocarbon dating The method was developed by Willard Libby in the late 1940s and soon became a standard tool for archaeologists. Libby received the Nobel Prize in Chemistry for his work in 1960. The radiocarbon dating method is based on the fact that radiocarbon is constantly being created in the atmosphere by the interaction of cosmic rays with atmospheric nitrogen. The resulting radiocarbon combines with atmospheric oxygen to form radioactive carbon dioxide, which is incorporated into plants by photosynthesis; animals then acquire 14C by eating the plants. When the animal or plant dies, it stops exchanging carbon with its environment, and from that point onwards the amount of 14C it contains begins to decrease as the 14C undergoes radioactive decay. Measuring the amount of 14C in a sample from a dead plant or animal such as a piece of wood or a fragment of bone provides information that can be used to calculate when the animal or plant died. The older a sample is, the less 14C there is to be detected, and because the half-life of 14C (the period of time after which half of a given sample will have decayed) is about 5,730 years, the oldest dates that can be reliably measured by this process date to around 50,000 years ago, although special preparation methods occasionally permit accurate analysis of older samples.