Caerorhachis
Caerorhachis (meaning "suitable spine" in Greek) is an extinct genus of early tetrapod from the Early Carboniferous of Scotland, probably from the Serpukhovian stage. Its placement within Tetrapoda is uncertain, but it is generally regarded as a primitive member of the group. The type species C. bairdi was named in 1977. Classification Caerorhachis has usually been placed as a basal anthracosaur or a close relative of anthracosaurs. In this classification, Caerorhachis is a close ancestor of amniotes, or tetrapods that lay eggs on land. Caerorhachis has also been classified as the sister taxon of temnospondyls, a large group of extinct amphibians, based on the presence of several primitive traits. In fact, when it was named in 1977, Caerorhachis was thought to be a dendrerpetontid temnospondyl. The vertebrae of Caerorhachis are more similar to anthracosaurs, however. As in all early tetrapods, the centrum, or central part of the vertebra, is composed of two parts: the intercentrum and the pleurocentrum. While temnospondyls have large intercentra and small pleurocentra, Caerorhachis and anthracosaurs have larger pleurocentra than intercentra. A 2003 phylogenetic analysis of early tetrapods placed Caerorhachis outside the clade that included temnospondyls and anthracosaurs in an ancestral position to both groups. Paleobiology Caerorhachis